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Paenungulata
Paenungulata (from Latin ''paene'' "almost" + ''ungulātus'' "having Hoof, hoofs") is a clade of "sub-Ungulate, ungulates", which groups three Extant taxon, extant mammal Order (biology), orders: Proboscidea (including elephants), Sirenia (sirenia, sea cows, including dugongs and manatees), and Hyracoidea (hyraxes). At least two more possible orders are known only as fossils, namely Embrithopoda and Desmostylia. Molecular evidence indicates that Paenungulata (or at least its extant members) is part of the cohort Afrotheria, an ancient assemblage of mainly African mammals of great diversity. The other members of this cohort are the orders Afrosoricida (tenrecs and golden moles), Macroscelidea (elephant shrews) and Tubulidentata (aardvarks). Of the five orders, hyraxes are the most basal, followed by embrithopods; the remaining orders (sirenians and elephants) are more closely related. These latter three are grouped as the Tethytheria, because it is believed that their common ancest ...
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Ocepeia
''Ocepeia'' is an extinct genus of afrotherian mammal that lived in present-day Morocco during the middle Paleocene epoch, approximately 60 million years ago. First named and described in 2001, the type species is ''O. daouiensis'' from the Selandian stage of Morocco's Ouled Abdoun Basin. A second, larger species, ''O. grandis'', is known from the Thanetian, a slightly younger stage in the same area. In life, the two species are estimated to have weighed about and , respectively, and are believed to have been specialized folivore, leaf-eaters. The fossil skulls of ''Ocepeia'' are the oldest known afrotherian skulls, and the best-known of any Paleocene mammal in Africa. Loosely grouped with the archaic ungulates known as Condylarth, "condylarths", ''Ocepeia'' shares several features with primitive Paenungulata, paenungulates (a group including elephants, hyraxes, and extinct relatives), but some analyses suggest it is more closely related to Afroinsectiphilia (a group containing ...
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Sirenia
The Sirenia (), commonly referred to as sea-cows or sirenians, are an order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The Sirenia currently comprise two distinct families: Dugongidae (the dugong and the now extinct Steller's sea cow) and Trichechidae (manatees, namely the Amazonian manatee, West Indian manatee, and West African manatee) with a total of four species. The Protosirenidae (Eocene sirenians) and Prorastomidae (terrestrial sirenians) families are extinct. Sirenians are classified in the clade Paenungulata, alongside the elephants and the hyraxes, and evolved in the Eocene 50 million years ago (mya). The Dugongidae diverged from the Trichechidae in the late Eocene or early Oligocene (30–35 mya). Sirenians grow to between in length and in weight. The historic Steller's sea cow was the largest known sirenian to have lived, and could reach lengths of and weights of . Sirenians have a ...
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Afrotheria
Afrotheria ( from Latin ''Afro-'' "of Africa" + ''theria'' "wild beast") is a clade of mammals, the living members of which belong to groups that are either currently living in Africa or of African origin: golden moles, elephant shrews (also known as sengis), tenrecs, aardvarks, hyraxes, elephants, sea cows, and several extinct clades. Most groups of afrotheres share little or no superficial resemblance, and their similarities have only become known in recent times because of genetics and molecular studies. Many afrothere groups are found mostly or exclusively in Africa, reflecting the fact that Africa was an island continent from the Cretaceous until the early Miocene around 20 million years ago, when Afro-Arabia collided with Eurasia. Because Africa was isolated by water, Laurasian groups of mammals such as insectivores, rodents, lagomorphs, carnivorans and ungulates could not reach Africa for much of the early to mid-Cenozoic. Instead, the niches occupied by those groups on the ...
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Sirenia
The Sirenia (), commonly referred to as sea-cows or sirenians, are an order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The Sirenia currently comprise two distinct families: Dugongidae (the dugong and the now extinct Steller's sea cow) and Trichechidae (manatees, namely the Amazonian manatee, West Indian manatee, and West African manatee) with a total of four species. The Protosirenidae (Eocene sirenians) and Prorastomidae (terrestrial sirenians) families are extinct. Sirenians are classified in the clade Paenungulata, alongside the elephants and the hyraxes, and evolved in the Eocene 50 million years ago (mya). The Dugongidae diverged from the Trichechidae in the late Eocene or early Oligocene (30–35 mya). Sirenians grow to between in length and in weight. The historic Steller's sea cow was the largest known sirenian to have lived, and could reach lengths of and weights of . Sirenians have a ...
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Desmostylia
The Desmostylia (from Greek δεσμά ''desma'', "bundle", and στῦλος ''stylos'', "pillar") are an extinct order of aquatic mammals that existed from the early Oligocene (Rupelian) to the late Miocene (Tortonian) (). Desmostylians are the only known extinct order of marine mammals. The Desmostylia, together with Sirenia and Proboscidea (and possibly Embrithopoda), have traditionally been assigned to the afrotherian clade Tethytheria, a group named after the paleoocean Tethys around which they originally evolved. The relationship between the Desmostylia and the other orders within the Tethytheria has been disputed; if the common ancestor of all tethytheres was semiaquatic, the Proboscidea became secondarily terrestrial; alternatively, the Desmostylia and Sirenia could have evolved independently into aquatic mammals. The assignment of Desmostylia to Afrotheria has always been problematic from a biogeographic standpoint, given that Africa was the locus of the early evoluti ...
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Hyracoidea
Hyraxes (), also called dassies, are small, thickset, herbivorous mammals in the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Typically, they measure between long and weigh between . They are superficially similar to pikas and marmots, but are more closely related to elephants and sea cows. Hyraxes have a life span from 9 to 14 years. Five extant species are recognised: the rock hyrax (''Procavia capensis'') and the yellow-spotted rock hyrax (''Heterohyrax brucei''), which both live on rock outcrops, including cliffs in Ethiopia and isolated granite outcrops called koppies in southern Africa; the western tree hyrax (''Dendrohyrax dorsalis''), southern tree hyrax (''D. arboreus''), and eastern tree hyrax (''D. validus''). Their distribution is limited to Africa, except for ''P. capensis'', which is also found in the Middle East. Characteristics Hyraxes retain or have redeveloped a number of primitive mammalian characteristics; in particular, they ...
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Hyrax
Hyraxes (), also called dassies, are small, thickset, herbivorous mammals in the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Typically, they measure between long and weigh between . They are superficially similar to pikas and marmots, but are more closely related to elephants and sea cows. Hyraxes have a life span from 9 to 14 years. Five extant species are recognised: the rock hyrax (''Procavia capensis'') and the yellow-spotted rock hyrax (''Heterohyrax brucei''), which both live on rock outcrops, including cliffs in Ethiopia and isolated granite outcrops called koppies in southern Africa; the western tree hyrax (''Dendrohyrax dorsalis''), southern tree hyrax (''D. arboreus''), and eastern tree hyrax (''D. validus''). Their distribution is limited to Africa, except for ''P. capensis'', which is also found in the Middle East. Characteristics Hyraxes retain or have redeveloped a number of primitive mammalian characteristics; in particular, the ...
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Elephant
Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae and the order Proboscidea. The order was formerly much more diverse during the Pleistocene, but most species became extinct during the Late Pleistocene epoch. Distinctive features of elephants include a long proboscis called a trunk, tusks, large ear flaps, pillar-like legs, and tough but sensitive skin. The trunk is used for breathing, bringing food and water to the mouth, and grasping objects. Tusks, which are derived from the incisor teeth, serve both as weapons and as tools for moving objects and digging. The large ear flaps assist in maintaining a constant body temperature as well as in communication. African elephants have larger ears and concave backs, whereas Asian elephants have smaller ears, and convex or level backs. Elephants ...
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Ungulate
Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. The term means, roughly, "being hoofed" or "hoofed animal". As a descriptive term, "ungulate" normally excludes cetaceans as they do not possess most of the typical morphological characteristics of other ungulates, but recent discoveries indicate that they were also descended from early artiodactyls. Ungulates are typically herbivorous and many employ specialized gut bacteria to allow them to digest cellulose. Some modern species, such as pigs, are omnivorous, ...
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Tethytheria
Tethytheria is a clade of mammals that includes the sirenians and proboscideans, as well as the extinct order Embrithopoda. Though there is strong anatomical and molecular support for the monophyly of Tethytheria, the interrelationships between the included taxa remain disputed. The tethytheres are united by several characters, including anteriorly facing orbits and more or less bilophodont cheek teeth (double transverse ridges on the crowns of the teeth). Proboscidea and Sirenia are linked together based on auditory characters in their petrosal bones, but this link may be a homoplasy. Desmostylians, traditionally considered tethytheres, have been tentatively assigned to Perissodactyla, along with the Early Eocene family Anthracobunidae, which was considered a sister group to Tethytheria. Systematics Cladogram modified from and . Classification Classification modified from . * Clade Tethytheria ** order †Embrithopoda *** † Phenacolophidae *** †Arsinoitheriidae ** order ...
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Embrithopoda
Embrithopoda ("heavy-footed") is an order of extinct mammals known from Asia, Africa and Eastern Europe. Most of the embrithopod genera are known exclusively from jaws and teeth dated from the late Paleocene to the late Eocene; however, the order is best known from its terminal member, the elephantine ''Arsinoitherium''. Description While embrithopods bore a superficial resemblance to rhinoceroses, their horns had bony cores covered in keratinized skin. Not all embrithopods possessed horns, either. Despite their appearance, they have been regarded as related to elephants, not perissodactyls. As tethytheres, the Embrithopoda have been believed to be part of the clade Afrotheria. However, a study of the basal arsinoitheriid, ''Palaeoamasia'', suggests that embrithopods are not tethytheres or even paenungulates, and that they need to be better sampled in an analysis of eutherian relationships to clarify if they are even afrotherians.[Erdal et al.'s inclusion of Embrithopoda in ...
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Arsinoitherium
''Arsinoitherium'' is an extinct genus of paenungulate mammals belonging to the extinct Order (biology), order Embrithopoda. It is related to elephants, sirenians, hyraxes and the extinct desmostylians. Arsinoitheres were superficially rhinoceros-like herbivores that lived during the Late Eocene and the Early Oligocene of North Africa from 36 to 30 million years ago, in areas of tropical rainforest and at the margin of mangrove swamps. A species described in 2004, ''A. giganteum'', lived in Ethiopia about 27 million years ago. Taxonomy The best-known (and first-described) species is ''A. zitteli''. Another species, ''A. giganteum'', was discovered in the Ethiopian highlands of Chilga in 2003. The fossil teeth, far larger than those of ''A. zitteli'', date to around 28–27 million years ago. While the Fayum Oasis is the only site where complete skeletons of ''Arsinoitherium'' fossils were recovered, arsinoitheriids have been found in southeastern Europe, including ''Crivadiather ...
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