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Embrithopoda
Embrithopoda ("heavy-footed") is an order of extinct mammals known from Asia, Africa and Eastern Europe. Most of the embrithopod genera are known exclusively from jaws and teeth dated from the late Paleocene to the late Eocene; however, the order is best known from its terminal member, the elephantine ''Arsinoitherium''. Description While embrithopods bore a superficial resemblance to rhinoceroses, their horns had bony cores covered in keratinized skin. Not all embrithopods possessed horns, either. Despite their appearance, they have been regarded as related to elephants, not perissodactyls. As tethytheres, the Embrithopoda have been believed to be part of the clade Afrotheria. However, a study of the basal arsinoitheriid, ''Palaeoamasia'', suggests that embrithopods are not tethytheres or even paenungulates, and that they need to be better sampled in an analysis of eutherian relationships to clarify if they are even afrotherians.[Erdal et al.'s inclusion of Embrithopoda in ...
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Paenungulata
Paenungulata (from Latin ''paene'' "almost" + ''ungulātus'' "having Hoof, hoofs") is a clade of "sub-Ungulate, ungulates", which groups three Extant taxon, extant mammal Order (biology), orders: Proboscidea (including elephants), Sirenia (sirenia, sea cows, including dugongs and manatees), and Hyracoidea (hyraxes). At least two more possible orders are known only as fossils, namely Embrithopoda and Desmostylia. Molecular evidence indicates that Paenungulata (or at least its extant members) is part of the cohort Afrotheria, an ancient assemblage of mainly African mammals of great diversity. The other members of this cohort are the orders Afrosoricida (tenrecs and golden moles), Macroscelidea (elephant shrews) and Tubulidentata (aardvarks). Of the five orders, hyraxes are the most basal, followed by embrithopods; the remaining orders (sirenians and elephants) are more closely related. These latter three are grouped as the Tethytheria, because it is believed that their common ancest ...
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Afrotheria
Afrotheria ( from Latin ''Afro-'' "of Africa" + ''theria'' "wild beast") is a clade of mammals, the living members of which belong to groups that are either currently living in Africa or of African origin: golden moles, elephant shrews (also known as sengis), tenrecs, aardvarks, hyraxes, elephants, sea cows, and several extinct clades. Most groups of afrotheres share little or no superficial resemblance, and their similarities have only become known in recent times because of genetics and molecular studies. Many afrothere groups are found mostly or exclusively in Africa, reflecting the fact that Africa was an island continent from the Cretaceous until the early Miocene around 20 million years ago, when Afro-Arabia collided with Eurasia. Because Africa was isolated by water, Laurasian groups of mammals such as insectivores, rodents, lagomorphs, carnivorans and ungulates could not reach Africa for much of the early to mid-Cenozoic. Instead, the niches occupied by those groups on the ...
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Afrotheria
Afrotheria ( from Latin ''Afro-'' "of Africa" + ''theria'' "wild beast") is a clade of mammals, the living members of which belong to groups that are either currently living in Africa or of African origin: golden moles, elephant shrews (also known as sengis), tenrecs, aardvarks, hyraxes, elephants, sea cows, and several extinct clades. Most groups of afrotheres share little or no superficial resemblance, and their similarities have only become known in recent times because of genetics and molecular studies. Many afrothere groups are found mostly or exclusively in Africa, reflecting the fact that Africa was an island continent from the Cretaceous until the early Miocene around 20 million years ago, when Afro-Arabia collided with Eurasia. Because Africa was isolated by water, Laurasian groups of mammals such as insectivores, rodents, lagomorphs, carnivorans and ungulates could not reach Africa for much of the early to mid-Cenozoic. Instead, the niches occupied by those groups on the ...
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Tethytheria
Tethytheria is a clade of mammals that includes the sirenians and proboscideans, as well as the extinct order Embrithopoda. Though there is strong anatomical and molecular support for the monophyly of Tethytheria, the interrelationships between the included taxa remain disputed. The tethytheres are united by several characters, including anteriorly facing orbits and more or less bilophodont cheek teeth (double transverse ridges on the crowns of the teeth). Proboscidea and Sirenia are linked together based on auditory characters in their petrosal bones, but this link may be a homoplasy. Desmostylians, traditionally considered tethytheres, have been tentatively assigned to Perissodactyla, along with the Early Eocene family Anthracobunidae, which was considered a sister group to Tethytheria. Systematics Cladogram modified from and . Classification Classification modified from . * Clade Tethytheria ** order †Embrithopoda *** † Phenacolophidae *** †Arsinoitheriidae ** order ...
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Arsinoitheriidae
Arsinoitheriidae is a family of hoofed mammals belonging to the extinct order Embrithopoda. Remains have been found in the Middle East, Africa, Asia and Romania. When alive, they would have borne a strong but superficial resemblance to modern rhinoceroses; however, they were not closely related to them (or any other perissodactyl), instead being more closely related to hyraxes, elephants, sirenians, and possibly desmostylians (as part of the superorder afrotheria). Fossil record The last genus, ''Arsinoitherium'', was first recovered from the Latest Eocene of the Fayum; it disappears from the fossil record altogether before the end of the Early Oligocene. Etymology The name honors the wife of Ptolemy II, Queen Arsinoe II of Egypt, as the first fossils of ''Arsinoitherium ''Arsinoitherium'' is an extinct genus of paenungulate mammals belonging to the extinct order Embrithopoda. It is related to elephants, sirenians, hyraxes and the extinct desmostylians. Arsinoitheres were ...
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Palaeoamasiidae
Palaeoamasiidae or Palaeoamasinae is an extinct taxon of embrithopod mammals that have been found in Romania and Anatolia where they lived on the shores of the Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents .... Classification * Palaeoamasiidae ** '' Hypsamasia'' ** '' Palaeoamasia'' ** '' Crivadiatherium'' Radulesco, Iliesco &Iliesco 1976 Notes References * * * * * * Embrithopods Eocene mammals Eocene first appearances Eocene extinctions Prehistoric mammal families {{paleo-afrotheria-stub ...
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Arsinoitherium
''Arsinoitherium'' is an extinct genus of paenungulate mammals belonging to the extinct Order (biology), order Embrithopoda. It is related to elephants, sirenians, hyraxes and the extinct desmostylians. Arsinoitheres were superficially rhinoceros-like herbivores that lived during the Late Eocene and the Early Oligocene of North Africa from 36 to 30 million years ago, in areas of tropical rainforest and at the margin of mangrove swamps. A species described in 2004, ''A. giganteum'', lived in Ethiopia about 27 million years ago. Taxonomy The best-known (and first-described) species is ''A. zitteli''. Another species, ''A. giganteum'', was discovered in the Ethiopian highlands of Chilga in 2003. The fossil teeth, far larger than those of ''A. zitteli'', date to around 28–27 million years ago. While the Fayum Oasis is the only site where complete skeletons of ''Arsinoitherium'' fossils were recovered, arsinoitheriids have been found in southeastern Europe, including ''Crivadiather ...
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Arsinoitherium Zitteli
''Arsinoitherium'' is an extinct genus of paenungulate mammals belonging to the extinct order Embrithopoda. It is related to elephants, sirenians, hyraxes and the extinct desmostylians. Arsinoitheres were superficially rhinoceros-like herbivores that lived during the Late Eocene and the Early Oligocene of North Africa from 36 to 30 million years ago, in areas of tropical rainforest and at the margin of mangrove swamps. A species described in 2004, ''A. giganteum'', lived in Ethiopia about 27 million years ago. Taxonomy The best-known (and first-described) species is ''A. zitteli''. Another species, ''A. giganteum'', was discovered in the Ethiopian highlands of Chilga in 2003. The fossil teeth, far larger than those of ''A. zitteli'', date to around 28–27 million years ago. While the Fayum Oasis is the only site where complete skeletons of ''Arsinoitherium'' fossils were recovered, arsinoitheriids have been found in southeastern Europe, including '' Crivadiatherium'' fro ...
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Altungulata
Altungulata or Pantomesaxonia (''sensu'' and later authors) is an invalid clade (mirorder) of ungulate mammals comprising the perissodactyls, hyracoids, and tethytheres (sirenians, proboscideans, and related extinct taxa). The name "Pantomesaxonia" was originally introduced by , a German zoologist and racial theorist. It was resurrected by by including sirenians and excluding South American ungulates, phenacodontids, and meniscotheriids from the original concept. The name "Altungulata", introduced by and revised by , was erected as an alternative because the updated concept of "Pantomesaxonia" was regarded too deviant from the original concept. Both names are still in use, and, to add to the confusion, various authors assign different ranks to the involved taxa. For example, according to , Phenacodonta (Phenacodontidae and Meniscotheriidae) and Pantomesaxonia (Sirenia, Desmostylia, Proboscidea, Hyracoidea, and Perissodactyla) are sister groups together making up the supe ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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Ethiopia
Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east and northeast, Kenya to the south, South Sudan to the west, and Sudan to the northwest. Ethiopia has a total area of . As of 2022, it is home to around 113.5 million inhabitants, making it the 13th-most populous country in the world and the 2nd-most populous in Africa after Nigeria. The national capital and largest city, Addis Ababa, lies several kilometres west of the East African Rift that splits the country into the African and Somali tectonic plates. Anatomically modern humans emerged from modern-day Ethiopia and set out to the Near East and elsewhere in the Middle Paleolithic period. Southwestern Ethiopia has been proposed as a possible homeland of the Afroasiatic langua ...
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