Oradectes
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Oradectes
''Oradectes'' (meaning "margin biter", from Latin ''ora'' (margin) and ''dectes'' (biter)) is an extinct genus of diadectid reptiliomorph. It is known from a single partial skeleton collected from the Early Permian Cutler Formation of Colorado in the United States. The type species, ''O. sanmiguelensis'', was originally named as a species of ''Diadectes'' in 1965. It was given its own genus in 2010. Description The only material belonging to ''Oradectes'' is a partial skeleton known as MCZ 2989. This skeleton includes a complete skull, neck vertebrae, a partial pectoral girdle, the right forelimb, and some ribs. The skull is robust with spade-shaped teeth lining the jaws. The front teeth of the lower jaw protrude outward. A ventral border of the Meckelian fenestra of the lower jaw formed entirely from the splenial bone is a distinguishing feature of ''Oradectes'' found in no other diadectid. Classification ''Oradectes sanmiguelensis'' was described by Lewis and Vaughn (1965) as ...
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Diadectid
Diadectidae is an extinct family of early tetrapods that lived in what is now North America and Europe during the Late Carboniferous and Early Permian in Asia during the Late Permian. They were the first herbivorous tetrapods, and also the first fully terrestrial animals to attain large sizes. Footprints indicate that diadectids walked with an erect posture. They were the first to exploit plant material in terrestrial food chains, making their appearance an important stage in both vertebrate evolution and the development of terrestrial ecosystems. The best known and largest representative of the family is ''Diadectes'', a heavily built animal that attained a maximum length of several metres. Several other genera and various fragmentary fossil remains are also known. Although well known genera like ''Diadectes'' first appear in the Late Pennsylvanian, fragmentary remains of possible diadectids are known from much earlier deposits, including a piece of lower jaw found in Missis ...
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Diadectids
Diadectidae is an extinct family (biology), family of early tetrapods that lived in what is now North America and Europe during the Late Carboniferous and Early Permian in Asia during the Late Permian. They were the first herbivore, herbivorous tetrapods, and also the first fully terrestrial animals to attain large sizes. Footprints indicate that diadectids walked with an erect posture. They were the first to exploit plant material in terrestrial food chains, making their appearance an important stage in both vertebrate evolution and the development of terrestrial ecosystems. The best known and largest representative of the family is ''Diadectes'', a heavily built animal that attained a maximum length of several metres. Several other genera and various fragmentary fossil remains are also known. Although well known genera like ''Diadectes'' first appear in the Late Pennsylvanian, fragmentary remains of possible diadectids are known from much earlier deposits, including a piece of l ...
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Diadectes
''Diadectes'' (meaning ''crosswise-biter'') is an extinct genus of large reptiliomorphs or synapsids that lived during the early Permian period (Artinskian-Kungurian stages of the Cisuralian epoch, between 290 and 272 million years ago). ''Diadectes'' was one of the first herbivorous tetrapods, and also one of the first fully terrestrial vertebrates to attain large size. Description ''Diadectes'' was a heavily built animal, up to long, with a thick-boned skull, heavy vertebrae and ribs, massive limb girdles, and short, robust limbs. The nature of the limbs and vertebrae clearly indicates a terrestrial animal. The rib cage was assumed to be barrel-shaped, but new fossils show the ribs were actually sticking out to the sides. Paleobiology It possesses some characteristics of reptilians and amphibians, combining a reptile-like skeleton with a more primitive, seymouriamorph-like skull. ''Diadectes'' has been classified as belonging to the sister group of the amniotes. Among ...
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Silvadectes
''Diadectes'' (meaning ''crosswise-biter'') is an extinct genus of large reptiliomorphs or synapsids that lived during the early Permian period (Artinskian-Kungurian stages of the Cisuralian epoch, between 290 and 272 million years ago). ''Diadectes'' was one of the first herbivorous tetrapods, and also one of the first fully terrestrial vertebrates to attain large size. Description ''Diadectes'' was a heavily built animal, up to long, with a thick-boned skull, heavy vertebrae and ribs, massive limb girdles, and short, robust limbs. The nature of the limbs and vertebrae clearly indicates a terrestrial animal. The rib cage was assumed to be barrel-shaped, but new fossils show the ribs were actually sticking out to the sides. Paleobiology It possesses some characteristics of reptilians and amphibians, combining a reptile-like skeleton with a more primitive, seymouriamorph-like skull. ''Diadectes'' has been classified as belonging to the sister group of the amniotes. Among i ...
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Early Permian
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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Permian Colorado
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their amphibia ...
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Cisuralian Tetrapods Of North America
The Cisuralian is the first series/epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan and dates between 298.9 ± 0.15 – 272.3 ± 0.5 Mya. The Cisuralian is often synonymous with the informal terms early Permian or lower Permian. It corresponds approximately with the Wolfcampian in southwestern North America. The series saw the appearance of beetles and flies and was a relatively stable warming period of about 21 million years. Name and background The Cisuralian is the first series or epoch of the Permian. The Cisuralian was preceded by the last Pennsylvanian epoch (Gzhelian) and is followed by the Permian Guadalupian Epoch. The name "Cisuralian" was proposed in 1982, and approved by the International Subcommission on Permian Stratigraphy in 1996. The Cisuralian Epoch is named after the western slopes of the Ural Mountain ...
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Diasparactus
''Diasparactus'' is an extinct genus of diadectid reptiliomorphs, a group quite closely related to the amniotes, and paralleling some of their features. Like all advanced diadectids, ''Diasparactus'' was a herbivore, though not as large as its more well known relative ''Diadectes''. In ''Diasparactus'', the spines of the dorsal vertebrae are higher than in other genera in the family.A Description of a nearly complete skeleton of ''Diasparactus zenos'' Case
Permo-Carboniferous Vertebrates from New Mexico, p17-35. Retrieved 2011-09-11.


History

In 1910, E. C. Ca ...
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Desmatodon
''Desmatodon'' is an extinct genus of diadectid reptiliomorph. With fossils found from the Kasimovian (Missourian) stage of the Late Carboniferous of Pennsylvania, Colorado, and New Mexico in the United States, ''Desmatodon'' is the oldest known diadectid. Two species are currently recognized: the type species ''D. hollandi'' and the species ''D. hesperis''. Description Remains of ''Desmatodon'' have been found from the Glenshaw Formation of Pennsylvania, the Sangre de Cristo Formation in Colorado, and the Cutler Formation The Cutler Formation or Cutler Group is a rock unit that is exposed across the U.S. states of Arizona, northwest New Mexico, southeast Utah and southwest Colorado. It was laid down in the Early Permian during the Wolfcampian epoch. Descript ... of New Mexico. The genus is known mostly from teeth and portions of skulls. The cheek teeth are robust and spade-shaped with several cusps on their surfaces. The two species can be distinguished by the distribu ...
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Orobates
''Orobates'' is an extinct genus of diadectid reptiliomorphs that lived during the Early Permian. Its fossilised remains were found in Germany. A combination of primitive and derived traits (i.e. autapomorphic and plesiomorphic) distinguish it from all other well-known members of Diadectidae, a family of herbivorous reptiliomorphs. It weighed about 4 kg and appears to have been part of an upland fauna, browsing on high fibre plants. Locomotion The trace fossil species ''Ichniotherium sphaerodactylum'', from Bromacker in Germany, has been attributed to ''Orobates''. A study in 2015 found that the genus was characterized by a long body and tail, with fairly short legs and a short skull compared to the more derived ''Diadectes''. This indicates ''Orobates'' was less specialised for long treks compared to ''Diadectes''. A three-dimensional digital reconstruction of the holotype specimen allowed further analysis of the postcranium. An analysis of the mobility of the hip joi ...
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Ambedus
''Ambedus'' is an extinct genus of possible diadectid reptiliomorph. Fossils have been found from the Early Permian Dunkard Group of Monroe County, Ohio, Monroe County, Ohio. The type species ''A. pusillus'' was named in 2004. The genus name comes from the Latin word ''ambedo'' meaning "to nibble", in reference to its herbivorous diet. The specific name ''pusillus'' means "tiny" in Latin. Description ''Ambedus'' is a small tetrapod, possible Diadectid, known only from maxillae and dentary bones. It is usually considered the most primitive diadectid because unlike other forms it had a small, shallow lower jaw and many simple, conical teeth. It was also smaller than the later, rather bulky diadectids. Later diadectids have deep jaws with few teeth and forward-projecting incisor, incisiforms at the tips of the jaws for consuming plant material, and a corresponding greater girth. David Berman's 2013 paper argued against the inclusion of Ambedus in Diadectidae, for several reasons. He ...
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