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Diadectes
''Diadectes'' (meaning ''crosswise-biter'') is an extinct genus of large reptiliomorphs or synapsids that lived during the early Permian period (Artinskian-Kungurian stages of the Cisuralian epoch, between 290 and 272 million years ago). ''Diadectes'' was one of the first herbivorous tetrapods, and also one of the first fully terrestrial vertebrates to attain large size. Description ''Diadectes'' was a heavily built animal, up to long, with a thick-boned skull, heavy vertebrae and ribs, massive limb girdles, and short, robust limbs. The nature of the limbs and vertebrae clearly indicates a terrestrial animal. The rib cage was assumed to be barrel-shaped, but new fossils show the ribs were actually sticking out to the sides. Paleobiology It possesses some characteristics of reptilians and amphibians, combining a reptile-like skeleton with a more primitive, seymouriamorph-like skull. ''Diadectes'' has been classified as belonging to the sister group of the amniotes. Among ...
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Diadectes Scale
''Diadectes'' (meaning ''crosswise-biter'') is an extinct genus of large reptiliomorphs or synapsids that lived during the early Permian period (Artinskian-Kungurian stages of the Cisuralian epoch, between 290 and 272 million years ago). ''Diadectes'' was one of the first herbivore, herbivorous tetrapods, and also one of the first fully terrestrial vertebrates to attain large size. Description ''Diadectes'' was a heavily built animal, up to long, with a thick-boned skull, heavy vertebrae and ribs, massive limb girdles, and short, robust limbs. The nature of the limbs and vertebrae clearly indicates a terrestrial animal. The rib cage was assumed to be barrel-shaped, but new fossils show the ribs were actually sticking out to the sides. Paleobiology It possesses some characteristics of reptilians and amphibians, combining a reptile-like skeleton with a more primitive, Seymouriamorpha, seymouriamorph-like skull. ''Diadectes'' has been classified as belonging to the sister grou ...
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Diadectes Phaseolinus AMNH 4684 Skull
''Diadectes'' (meaning ''crosswise-biter'') is an extinct genus of large reptiliomorphs or synapsids that lived during the early Permian period (Artinskian-Kungurian stages of the Cisuralian epoch, between 290 and 272 million years ago). ''Diadectes'' was one of the first herbivorous tetrapods, and also one of the first fully terrestrial vertebrates to attain large size. Description ''Diadectes'' was a heavily built animal, up to long, with a thick-boned skull, heavy vertebrae and ribs, massive limb girdles, and short, robust limbs. The nature of the limbs and vertebrae clearly indicates a terrestrial animal. The rib cage was assumed to be barrel-shaped, but new fossils show the ribs were actually sticking out to the sides. Paleobiology It possesses some characteristics of reptilians and amphibians, combining a reptile-like skeleton with a more primitive, seymouriamorph-like skull. ''Diadectes'' has been classified as belonging to the sister group of the amniotes. Among i ...
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Silvadectes
''Diadectes'' (meaning ''crosswise-biter'') is an extinct genus of large reptiliomorphs or synapsids that lived during the early Permian period (Artinskian-Kungurian stages of the Cisuralian epoch, between 290 and 272 million years ago). ''Diadectes'' was one of the first herbivorous tetrapods, and also one of the first fully terrestrial vertebrates to attain large size. Description ''Diadectes'' was a heavily built animal, up to long, with a thick-boned skull, heavy vertebrae and ribs, massive limb girdles, and short, robust limbs. The nature of the limbs and vertebrae clearly indicates a terrestrial animal. The rib cage was assumed to be barrel-shaped, but new fossils show the ribs were actually sticking out to the sides. Paleobiology It possesses some characteristics of reptilians and amphibians, combining a reptile-like skeleton with a more primitive, seymouriamorph-like skull. ''Diadectes'' has been classified as belonging to the sister group of the amniotes. Among i ...
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Diadectes Tenuitecus
''Diadectes'' (meaning ''crosswise-biter'') is an extinct genus of large reptiliomorphs or synapsids that lived during the early Permian period (Artinskian-Kungurian stages of the Cisuralian epoch, between 290 and 272 million years ago). ''Diadectes'' was one of the first herbivorous tetrapods, and also one of the first fully terrestrial vertebrates to attain large size. Description ''Diadectes'' was a heavily built animal, up to long, with a thick-boned skull, heavy vertebrae and ribs, massive limb girdles, and short, robust limbs. The nature of the limbs and vertebrae clearly indicates a terrestrial animal. The rib cage was assumed to be barrel-shaped, but new fossils show the ribs were actually sticking out to the sides. Paleobiology It possesses some characteristics of reptilians and amphibians, combining a reptile-like skeleton with a more primitive, seymouriamorph-like skull. ''Diadectes'' has been classified as belonging to the sister group of the amniotes. Among i ...
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Tetrapod
Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids and all extant mammals). Tetrapods evolved from a clade of primitive semiaquatic animals known as the Tetrapodomorpha which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian period; their forms were transitional between lobe-finned fishes and true four-limbed tetrapods. Limbed vertebrates (tetrapods in the broad sense of the word) are first known from Middle Devonian trackways, and body fossils became common near the end of the Late Devonian but these were all aquatic. The first crown-tetrapods (last common ancestors of extant tetrapods capable of terrestrial locomotion) appeared by the very early Carboniferous, 350 million years ago. The specific aquatic ancestors ...
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Labyrinthodontia
"Labyrinthodontia" (Greek, 'maze-toothed') is an informal grouping of extinct predatory amphibians which were major components of ecosystems in the late Paleozoic and early Mesozoic eras (about 390 to 150 million years ago). Traditionally considered a subclass of the class Amphibia, modern classification systems recognize that labyrinthodonts are not a formal natural group (clade) exclusive of other tetrapods. Instead, they consistute an evolutionary grade (a paraphyletic group), ancestral to living tetrapods such as lissamphibians (modern amphibians) and amniotes (reptiles, mammals, and kin). "Labyrinthodont"-grade vertebrates evolved from lobe-finned fishes in the Devonian, though a formal boundary between fish and amphibian is difficult to define at this point in time. "Labyrinthodont" generally refers to extinct four-limbed tetrapods with a large body size and a crocodile-like lifestyle. The name describes the pattern of infolding of the dentin and enamel of the teeth, ...
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Cisuralian
The Cisuralian is the first Series (stratigraphy), series/Epoch (geology), epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian (geology), Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan and dates between 298.9 ± 0.15 – 272.3 ± 0.5 Mya. The Cisuralian is often synonymous with the informal terms early Permian or lower Permian. It corresponds approximately with the Wolfcampian in southwestern North America. The series saw the appearance of beetles and Fly, flies and was a relatively stable warming period of about 21 million years. Name and background The Cisuralian is the first series or epoch of the Permian. The Cisuralian was preceded by the last Pennsylvanian epoch (Gzhelian) and is followed by the Permian Guadalupian Epoch. The name "Cisuralian" was proposed in 1982, and approved by the International Subcommission on Permian Stratigraphy in 1996. The Ci ...
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Amniote
Amniotes are a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds, and extinct parareptiles and non-avian dinosaurs) and synapsids (including pelycosaurs and therapsids such as mammals). They are distinguished from the other tetrapod clade — the amphibians — by the development of three extraembryonic membranes ( amnion for embryoic protection, chorion for gas exchange, and allantois for metabolic waste disposal or storage), thicker and more keratinized skin, and costal respiration (breathing by expanding/constricting the rib cage). All three main features listed above, namely the presence of an amniotic buffer, water-impermeable cutes and a robust respiratory system, are very important for amniotes to live on land as true terrestrial animals – the ability to reproduce in locations away from water bodies, better homeostasis in drier environments, and more efficient air respiration to power terrestrial locomotions, although the ...
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Reptiliomorph
Reptiliomorpha (meaning reptile-shaped; in PhyloCode known as ''Pan-Amniota'') is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians ( lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes ''Homo sapiens'', but not ''Ascaphus truei'' (tailed frog). Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing ''Homo sapiens'', but not ''Pipa pipa'', '' Caecilia tentaculata'', and '' Siren lacertina''. The informal variant of the name, "reptiliomorphs", is also occasionally used to refer to stem-amniotes, i.e. a grade of reptile-like tetrapods that are more closely related to amniotes than they are to lissamphibians, but are not amniotes themselves; the name is used in this meaning e.g. by Ruta, Coates and Quicke (2003). An alternative name, "Anthracosauria", is also commonly used for the group, but is confusing ...
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Otic Notch
Otic notches are invaginations in the posterior margin of the skull roof, one behind each orbit. Otic notches are one of the features lost in the evolution of amniotes from their tetrapod ancestors. The notches have been interpreted as part of an auditory structure and are often reconstructed as holding a tympanum similar to those seen in modern anurans. Analysis of the ''columella'' (the stapes in amphibians and reptiles) of labyrinthodonts however indicates that it did not function in transmitting low-energy vibrations, thus rendering these animals effectively deaf to airborne sound. The otic notch instead functioned as a spiracle, at least in the early forms.Laurin, M. (1996)Hearing in Stegocephalians from the Tree of Life Web Project The Tree of Life Web Project is an Internet project providing information about the diversity and phylogeny of life on Earth. This collaborative peer reviewed project began in 1995, and is written by biologists from around the world. The site ...
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Artinskian
In the geologic timescale, the Artinskian is an age or stage of the Permian. It is a subdivision of the Cisuralian Epoch or Series. The Artinskian likely lasted between and million years ago (Ma) according to the most recent revision of the International Commission on Stratigraphy (ICS) in 2022. It was preceded by the Sakmarian and followed by the Kungurian. Stratigraphy The Artinskian is named after the small Russian city of Arti (formerly ''Artinsk''), situated in the southern Ural mountains, about 200 km southwest of Yekaterinburg. The stage was introduced into scientific literature by Alexander Karpinsky in 1874. Base of the Artinskian The base of the Artinskian Stage is defined as the first appearance datum (FAD) of the conodont species '' Sweetognathus whitei'' and ''Mesogondolella bisselli''. In order to constrain this age, the ICS subcommission on Permian stratigraphy informally proposed a candidate GSSP in 2002, later followed by a formal proposal in 201 ...
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