Natovenator Holotype Block
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Natovenator Holotype Block
''Natovenator'' is a genus of halszkaraptorine dromaeosaurid dinosaur from the Late Cretaceous Barun Goyot Formation of Mongolia. The genus is known from a single species, '' N. polydontus''. ''Natovenator'' is crucial to the understanding of halszkaraptorines due to it providing more support for the semi-aquatic lifestyle that has been proposed for this clade. This discovery is important as the semi-aquatic lifestyles of halszkaraptorines (mainly ''Halszkaraptor'') was contested in early 2022. Discovery and naming The ''Natovenator'' holotype specimen, MPC-D 102/114, was found in sediments of the Barun Goyot Formation of Omnogovi Province, Mongolia. It consists of a mostly articulated skeleton with a nearly complete skull. In 2022, ''Natovenator polydontus'' was described as a new genus and species of halszkaraptorine theropod dinosaurs by Sungjin Lee, Yuong-Nam Lee, Philip J. Currie, Robin Sissons, Jin-Young Park, Su-Hwan Kim, Rinchen Barsbold, and Khishigjav Tsogtbaatar ...
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Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary extinction event) occurred at the end of this age. In this mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous. Stratigraphic definitions Definition The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying rock strata of the Chalk Group c ...
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Piscivorous
A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted reptiles and synapsids evolved herbivory. Almost all predatory fishes (most sharks, tuna, billfishes, pikes etc.) are obligated piscivores. Some non-piscine aquatic animals, such as whales, sea lion and crocodilians, are not completely piscivorous, often also preying on invertebrates, marine mammals, waterbirds and even wading land animals in addition to fish; while others, such as the bulldog bat and gharial, are strictly dependent on fish for food. Some creatures, including cnidarians, octopuses, squid, spiders, cetaceans, grizzly bears, jaguars, wolves, snakes, turtles and sea gulls, may have fish as significa ...
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Hulsanpes
''Hulsanpes'' ( meaning " Khulsan foot") is a genus of halszkaraptorine theropod dinosaurs that lived during the Late Cretaceous in what is now the Barun Goyot Formation of Mongolia, about 75-72 million years ago. The remains were found in 1970 and formally described in 1982 by Halszka Osmólska, who noted that the genus is represented by an immature individual. ''Hulsanpes'' represents the first record of the basal dromaeosaurid subfamily Halszkaraptorinae. History of discovery The first fossil remains of ''Hulsanpes'' were discovered in 1970 during the third Polish-Mongolian expedition at Khulsan in the Gobi Desert. Twelve years later, the type species, ''Hulsanpes perlei'', was named and described by the Polish palaeontologist Halszka Osmólska in 1982. ''Hulsanpes'' is based on the holotype fossil specimen ZPAL MgD-I/173, uncovered in a sandstone layer of the Barun Goyot Formation, dating from the Late Campanian (roughly 72 million years ago). It consists of a partial brain ...
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Halszkaraptor Reconstruction By Tom Parker
''Halszkaraptor'' (; meaning " Halszka's seizer") is a genus of waterfowl-like dromaeosaurid dinosaurs from Mongolia that lived during the Late Cretaceous period. It contains only one known species, ''Halszkaraptor escuilliei''. The type specimen (holotype) has been compared to the bones of extant crocodilians and aquatic birds, and found evidence of a semiaquatic lifestyle, while some researchers question about semiaquatic ecology. A phylogenetic analysis revealed it was a member of the basal subfamily Halszkaraptorinae, along with ''Mahakala'' and ''Hulsanpes''. History of discovery The holotype specimen of ''Halszkaraptor'' likely came from the Djadochta Formation at Ukhaa Tolgod in southern Mongolia, and was illegally removed by fossil poachers. The fossil found its way to Japan and Great Britain, being owned by several collectors for some years until the Eldonia company of fossil dealer François Escuillié obtained it. He identified it as a new species, and in 2015 took ...
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Dromaeosauridae
Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning 'runner', and ('), meaning 'lizard'. In informal usage, they are often called raptors (after ''Velociraptor''), a term popularized by the film '' Jurassic Park''; a few types include the term "raptor" directly in their name and have come to emphasize their bird-like appearance and speculated bird-like behavior. Dromaeosaurid fossils have been found across the globe in North America, Europe, Africa, Asia, South America and Antarctica, with some fossils giving credence to the possibility that they inhabited Australia as well. They first appeared in the mid-Jurassic Period (late Bathonian stage, about 167 million years ago) and survived until the end of the Cretaceous (Maastrichtian stage, 66 ma), existing until the Cretaceous–Paleogene ex ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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Halszkaraptorinae
Halszkaraptorinae is a basal ("primitive") subfamily of Dromaeosauridae (or possibly Unenlagiidae) that includes the enigmatic genera ''Halszkaraptor'', ''Natovenator'', ''Mahakala'', and ''Hulsanpes''. Halszkaraptorines are definitively known only from Late Cretaceous strata in Asia, specifically in Mongolia. Following the recent discovery of ''Natovenator'', a member of the subfamily, the group is confirmed to have a semiaquatic lifestyle. History of discovery The first known remains of halszkaraptorines were found in sandstone sediments in 1970 during a Polish-Mongolian expedition at the Barun Goyot Formation, Gobi Desert. Later in 1982 they were described by Polish palaeontologist Halszka Osmólska and used as the holotype for the new genus and species '' Hulsanpes perlei'', honoring the Mongolian paleontologist Altangerel Perle. Although the affinities of these fossils were not fully understood, they were tentatively interpreted as belonging to Deinonychosauria. In 1992 dro ...
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Phylogenetics
In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλον, φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, Protein, protein Amino acid, amino acid sequences, or Morphology (biology), morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An un ...
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Natovenator Holotype Postcrania
''Natovenator'' is a genus of halszkaraptorine dromaeosaurid dinosaur from the Late Cretaceous Barun Goyot Formation of Mongolia. The genus is known from a single species, '' N. polydontus''. ''Natovenator'' is crucial to the understanding of halszkaraptorines due to it providing more support for the semi-aquatic lifestyle that has been proposed for this clade. This discovery is important as the semi-aquatic lifestyles of halszkaraptorines (mainly ''Halszkaraptor'') was contested in early 2022. Discovery and naming The ''Natovenator'' holotype specimen, MPC-D 102/114, was found in sediments of the Barun Goyot Formation of Omnogovi Province, Mongolia. It consists of a mostly articulated skeleton with a nearly complete skull. In 2022, ''Natovenator polydontus'' was described as a new genus and species of halszkaraptorine theropod dinosaurs by Sungjin Lee, Yuong-Nam Lee, Philip J. Currie, Robin Sissons, Jin-Young Park, Su-Hwan Kim, Rinchen Barsbold, and Khishigjav Tsogtbaatar ...
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Hesperornithiformes
Hesperornithes is an extinct and highly specialized group of aquatic avialans closely related to the ancestors of modern birds. They inhabited both marine and freshwater habitats in the Northern Hemisphere, and include genera such as ''Hesperornis'', ''Parahesperornis'', ''Baptornis'', ''Enaliornis'', and ''Potamornis'', all strong-swimming, predatory divers. Many of the species most specialized for swimming were completely flightless. The largest known hesperornithean, ''Canadaga arctica'', may have reached a maximum adult length of . Hesperornitheans were the only Mesozoic avialans to colonize the oceans. They were wiped out in the Cretaceous–Paleogene extinction event, along with enantiornitheans and all other non-avian dinosaurs, and many other diverse plant and animal groups. Anatomy and ecology Most of what is known about this group rests on analyses of single species, as few provide sufficiently complete fossils for analysis. Although some of the smaller and more basal ...
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Premaxilla
The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has been usually termed as the incisive bone. Other terms used for this structure include premaxillary bone or ''os premaxillare'', intermaxillary bone or ''os intermaxillare'', and Goethe's bone. Human anatomy In human anatomy, the premaxilla is referred to as the incisive bone (') and is the part of the maxilla which bears the incisor teeth, and encompasses the anterior nasal spine and alar region. In the nasal cavity, the premaxillary element projects higher than the maxillary element behind. The palatal portion of the premaxilla is a bony plate with a generally transverse orientation. The incisive foramen is bound anteriorly and laterally by the premaxilla and posteriorly by the palatine process of the maxilla. It is formed from the ...
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Waterfowl
Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which includes over 170 species of waterfowl, among them the ducks, geese, and swans. Most modern species in the order are highly adapted for an aquatic existence at the water surface. With the exception of screamers, males have penises, a trait that has been lost in the Neoaves. Due to their aquatic nature, most species are web-footed. Evolution Anseriformes are one of only two types of modern bird to be confirmed present during the Mesozoic alongside the other dinosaurs, and in fact were among the very few birds to survive their extinction, along with their cousins the galliformes. These two groups only occupied two ecological niches during the Mesozoic, living in water and on the ground, while the toothed enantiornithes were the dominant bird ...
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