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Monocot
Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of the major groups into which the flowering plants have traditionally been divided; the rest of the flowering plants have two cotyledons and are classified as dicotyledons, or dicots. Monocotyledons have almost always been recognized as a group, but with various taxonomic ranks and under several different names. The APG III system of 2009 recognises a clade called "monocots" but does not assign it to a taxonomic rank. The monocotyledons include about 60,000 species, about a quarter of all angiosperms. The largest family in this group (and in the flowering plants as a whole) by number of species are the orchids (family Orchidaceae), with more than 20,000 species. About half as many species belong to the true grasses ( Poaceae), which are ...
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Lilioid Monocots
Lilioid monocots (lilioids, liliid monocots, petaloid monocots, petaloid lilioid monocots) is an informal name used for a grade (grouping of taxa with common characteristics) of five monocot orders (Petrosaviales, Dioscoreales, Pandanales, Liliales and Asparagales) in which the majority of species have flowers with relatively large, coloured tepals. This characteristic is similar to that found in lilies ("lily-like"). Petaloid monocots refers to the flowers having tepals which all resemble petals (petaloid). The taxonomic terms Lilianae or Liliiflorae have also been applied to this assemblage at various times. From the early nineteenth century many of the species in this group of plants were put into a very broadly defined family, Liliaceae ''sensu lato'' or ''s.l.'' (lily family). These classification systems are still found in many books and other sources. Within the monocots the Liliaceae ''s.l.'' were distinguished from the Glumaceae. The development of molecular phyloge ...
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Asparagales
Asparagales (asparagoid lilies) is an order (biology), order of plants in modern classification systems such as the Angiosperm Phylogeny Group (APG) and the Angiosperm Phylogeny Web. The order takes its name from the type (biology), type family (biology), family Asparagaceae and is placed in the monocots amongst the lilioid monocots. The order has only recently been recognized in classification systems. It was first put forward by herbert Huber (botanist), Huber in 1977 and later taken up in the Dahlgren system of 1985 and then the APG in 1998, 2003 and 2009. Before this, many of its families were assigned to the old order Liliales, a very large order containing almost all monocots with colorful tepals and lacking starch in their endosperm. DNA sequencing, DNA sequence analysis indicated that many of the taxa previously included in Liliales should actually be redistributed over three orders, Liliales, Asparagales, and Dioscoreales. The boundaries of the Asparagales and of its fam ...
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Lilianae
Lilianae (also known as Liliiflorae) is a botanical name for a superorder (that is, a rank higher than that of order) of flowering plants. Such a superorder of necessity includes the type family Liliaceae (and usually the type order Liliales). Terminations at the rank of superorder are not standardized by the International Code of Nomenclature for algae, fungi, and plants (ICN), although the suffix ''-anae'' has been proposed. Lilianae, introduced in 1966 as a name for a superorder, progressively replaced the older term Liliiflorae, introduced in 1825 as a name for an order. Taxonomy History Early history - Liliiflorae Liliiflorae was a term introduced by Carl Adolph Agardh in 1825 as a higher order to include the Liliaceae (which he called Coronariae) and related families. Argadh, together with De Candolle developed the concept of ordered botanical ranks, in this case grouping together De Jussieu's (1789) recently defined collections of genera (families) into high ...
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Liliales
Liliales is an order of monocotyledonous flowering plants in the Angiosperm Phylogeny Group and Angiosperm Phylogeny Web system, within the lilioid monocots. This order of necessity includes the family Liliaceae. The APG III system (2009) places this order in the monocot clade. In APG III, the family Luzuriagaceae is combined with the family Alstroemeriaceae and the family Petermanniaceae is recognized. Both the order Lililiales and the family Liliaceae have had a widely disputed history, with the circumscription varying greatly from one taxonomist to another. Previous members of this order, which at one stage included most monocots with conspicuous tepals and lacking starch in the endosperm are now distributed over three orders, Liliales, Dioscoreales and Asparagales, using predominantly molecular phylogenetics. The newly delimited Liliales is monophyletic, with ten families. Well known plants from the order include ''Lilium'' (lily), tulip, the North American wildflower ''Tri ...
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Zingiberales
The Zingiberales are flowering plants forming one of four orders in the commelinids clade of monocots, together with its sister order, Commelinales. The order includes 68 genera and 2,600 species. Zingiberales are a unique though morphologically diverse order that has been widely recognised as such over a long period of time. They are usually large herbaceous plants with rhizomatous root systems and lacking an aerial stem except when flowering. Flowers are usually large and showy, and the stamens are often modified ( staminodes) to also form colourful petal-like structures that attract pollinators. Zingiberales contain eight families that are informally considered as two groups, differing in the number of fertile stamens. A " banana group" of four families appeared first and were named on the basis of large banana-like leaves. Later, a more genetically coherent (monophyletic) " ginger group" appeared, consisting of the remaining four families. The order, which has a fossil re ...
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Dioscoreales
The Dioscoreales are an order of monocotyledonous flowering plants in modern classification systems, such as the Angiosperm Phylogeny Group and the Angiosperm Phylogeny Web. Within the monocots Dioscoreales are grouped in the lilioid monocots where they are in a sister group relationship with the Pandanales. The Dioscoreales must contain the family Dioscoreaceae which includes the yam ('' Dioscorea'') which is an important food source in many regions. Older systems tended to place all lilioid monocots with reticulate veined leaves (such as Smilacaceae and Stemonaceae together with Dioscoraceae) in Dioscoreales. As currently circumscribed by phylogenetic analysis using combined morphology and molecular methods, Dioscreales contains many reticulate veined vines in Dioscoraceae, it also includes the myco-heterotrophic Burmanniaceae and the autotrophic Nartheciaceae. The order consists of three families, 22 genera and about 850 species. Description Dioscoreales are vines or her ...
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Alismatid Monocots
Alismatid monocots (alismatids, basal monocots) is an informal name for a group of early branching (hence basal) monocots, consisting of two orders, the Acorales and Alismatales. The name has also been used to refer to the Alismatales alone. Monocots are frequently treated as three informal groupings based on their branching from ancestral monocots and shared characteristics: alismatid monocots, lilioid monocots (the five other non-commelinid monocots) and commelinid monocots. Research at the Royal Botanical Gardens, Kew is organised into two teams I: Alismatids and Lilioids and II: Commelinids. A similar approach is taken by Judd in his ''Plant systematics''. Phylogeny Cladogram showing the orders of monocots (Lilianae ''sensu'' Chase & Reveal) based on molecular phylogenetic evidence according to the Angiosperm Phylogeny Group IV (APG IV). Subdivision Of the two orders, the Acorales is monotypic, consisting of a single family, the Acoraceae, which in turn has ...
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Liliopsida
Liliopsida Batsch (synonym: Liliatae) is a botanical name for the class containing the family Liliaceae (or Lily Family). It is considered synonymous (or nearly synonymous) with the name monocotyledon. Publication of the name is credited to Scopoli (in 1760): see author citation (botany). This name is formed by replacing the termination ''-aceae'' in the name Liliaceae by the termination ''-opsida'' (Art 16 of the ICBN). Although in principle it is true that circumscription of this class will vary with the taxonomic system being used, in practice this name is very strongly linked to the Cronquist system, and the allied Takhtajan system. These two are the only major systems to use the name, and in both these systems it refers to the group more widely known as the monocotyledons. Earlier systems referred to this group by the name Monocotyledones, with Monocotyledoneae an earlier spelling (these names may be used in any rank). Systems such as the Dahlgren and Thorne systems (more ...
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Commelinid
In plant taxonomy, commelinids (originally commelinoids) (plural, not capitalised) is a clade of flowering plants within the monocots, distinguished by having cell walls containing ferulic acid. The commelinids are the only clade that the APG IV system has informally named within the monocots. The remaining monocots are a paraphyletic unit. Also known as the commelinid monocots it forms one of three groupings within the monocots, and the final branch; the other two groups are the alismatid monocots and the lilioid monocots. Description Members of the commelinid clade have cell walls containing UV-fluorescent ferulic acid. Taxonomy The commelinids were first recognized as a formal group in 1967 by Armen Takhtajan, who named them the Commelinidae and assigned them to a subclass of Liliopsida (monocots). The name was also used in the 1981 Cronquist system. However, by the release of his 1980 system of classification, Takhtajan had merged this subclass into a larger one, and no ...
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Acorales
''Acorus'' is a genus of monocot flowering plants. This genus was once placed within the family Araceae (aroids), but more recent classifications place it in its own family Acoraceae and order Acorales, of which it is the sole genus of the oldest surviving line of monocots. Some older studies indicated that it was placed in a lineage (the order Alismatales), that also includes aroids (Araceae), Tofieldiaceae, and several families of aquatic monocots (e.g., Alismataceae, Posidoniaceae). However, modern phylogenetic studies demonstrate that ''Acorus'' is sister to all other monocots. Common names include calamus and sweet flag. The genus is native to North America and northern and eastern Asia, and naturalised in southern Asia and Europe from ancient cultivation. The known wild populations are diploid except for some tetraploids in eastern Asia, while the cultivated plants are sterile triploids, probably of hybrid origin between the diploid and tetraploid forms. Characteristics ...
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Alismatales
The Alismatales (alismatids) are an order of flowering plants including about 4,500 species. Plants assigned to this order are mostly tropical or aquatic. Some grow in fresh water, some in marine habitats. Description The Alismatales comprise herbaceous flowering plants of often aquatic and marshy habitats, and the only monocots known to have green embryos other than the Amaryllidaceae. They also include the only marine angiosperms growing completely submerged, the seagrasses. The flowers are usually arranged in inflorescences, and the mature seeds lack endosperm. Both marine and freshwater forms include those with staminate flowers that detach from the parent plant and float to the surface. There they can pollinate carpellate flowers floating on the surface via long pedicels. In others, pollination occurs underwater, where pollen may form elongated strands, increasing chance of success. Most aquatic species have a totally submerged juvenile phase, and flowers are either ...
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Pandanales
Pandanales, the pandans or screw-pines, is an order of flowering plants placed in the monocot clade in the Angiosperm Phylogeny Group and Angiosperm Phylogeny Web systems. Within the monocots Pandanales are grouped in the lilioid monocots where they are in a sister group relationship with the Dioscoreales. Historically the order has consisted of a number of different families in different systems but modern classification of the order is based primarily on molecular phylogenetics despite diverse morphology which previously placed many of the families in other groupings based on apparent similarity. Members of the order have a subtropical distribution and includes trees, shrubs, and vines as well as herbaceous plants. The order consists of 5 families, 36 genera and about 1,610 species. Description Pandanales are highly diverse including large arboraceous plants of tropical rainforests and coastal areas, climbing vines and lianas, as well as very small achlorophyllous (mycoh ...
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