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Liliales
Liliales is an order of monocotyledonous flowering plants in the Angiosperm Phylogeny Group and Angiosperm Phylogeny Web system, within the lilioid monocots. This order of necessity includes the family Liliaceae. The APG III system (2009) places this order in the monocot clade. In APG III, the family Luzuriagaceae is combined with the family Alstroemeriaceae and the family Petermanniaceae is recognized. Both the order Lililiales and the family Liliaceae have had a widely disputed history, with the circumscription varying greatly from one taxonomist to another. Previous members of this order, which at one stage included most monocots with conspicuous tepals and lacking starch in the endosperm are now distributed over three orders, Liliales, Dioscoreales and Asparagales, using predominantly molecular phylogenetics. The newly delimited Liliales is monophyletic, with ten families. Well known plants from the order include ''Lilium'' (lily), tulip, the North American wildflower ''Tri ...
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Lilioid Monocots
Lilioid monocots (lilioids, liliid monocots, petaloid monocots, petaloid lilioid monocots) is an informal name used for a grade (grouping of taxa with common characteristics) of five monocot orders (Petrosaviales, Dioscoreales, Pandanales, Liliales and Asparagales) in which the majority of species have flowers with relatively large, coloured tepals. This characteristic is similar to that found in lilies ("lily-like"). Petaloid monocots refers to the flowers having tepals which all resemble petals (petaloid). The taxonomic terms Lilianae or Liliiflorae have also been applied to this assemblage at various times. From the early nineteenth century many of the species in this group of plants were put into a very broadly defined family, Liliaceae ''sensu lato'' or ''s.l.'' (lily family). These classification systems are still found in many books and other sources. Within the monocots the Liliaceae ''s.l.'' were distinguished from the Glumaceae. The development of molecular phyloge ...
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Liliaceae
The lily family, Liliaceae, consists of about 15 genera and 610 species of flowering plants within the order Liliales. They are monocotyledonous, perennial, herbaceous, often bulbous geophytes. Plants in this family have evolved with a fair amount of morphological diversity despite genetic similarity. Common characteristics include large flowers with parts arranged in threes: with six colored or patterned petaloid tepals (undifferentiated petals and sepals) arranged in two whorls, six stamens and a superior ovary. The leaves are linear in shape, with their veins usually arranged parallel to the edges, single and arranged alternating on the stem, or in a rosette at the base. Most species are grown from bulbs, although some have rhizomes. First described in 1789, the lily family became a paraphyletic "catch-all" (wastebasket) group of lilioid monocots that did not fit into other families and included a great number of genera now included in other families and in some cases in ...
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Monocotyledon
Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of the major groups into which the flowering plants have traditionally been divided; the rest of the flowering plants have two cotyledons and are classified as dicotyledons, or dicots. Monocotyledons have almost always been recognized as a group, but with various taxonomic ranks and under several different names. The APG III system of 2009 recognises a clade called "monocots" but does not assign it to a taxonomic rank. The monocotyledons include about 60,000 species, about a quarter of all angiosperms. The largest family in this group (and in the flowering plants as a whole) by number of species are the orchids (family Orchidaceae), with more than 20,000 species. About half as many species belong to the true grasses ( Poaceae), which are ...
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Monocot
Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of the major groups into which the flowering plants have traditionally been divided; the rest of the flowering plants have two cotyledons and are classified as dicotyledons, or dicots. Monocotyledons have almost always been recognized as a group, but with various taxonomic ranks and under several different names. The APG III system of 2009 recognises a clade called "monocots" but does not assign it to a taxonomic rank. The monocotyledons include about 60,000 species, about a quarter of all angiosperms. The largest family in this group (and in the flowering plants as a whole) by number of species are the orchids (family Orchidaceae), with more than 20,000 species. About half as many species belong to the true grasses ( Poaceae), which are ...
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Campynemataceae
Campynemataceae (Campynemaceae) is a family of flowering plants. The family consists of two genera and four species of perennial herbaceous plants endemic to New Caledonia and Tasmania. Taxonomy Originally described by Dumortier in 1829, Campynemaceae consisted of a single genus, '' Campynema'', described by Labillardière in Tasmania in 1804. In 1893 Baillon identified a closely related genus, '' Campynemanthe'' in New Caledonia. Together the two genera make up the family Campynemataceae sensu Angiosperm Phylogeny Group (APG), within the Liliales order. While historically, the two genera have generally been treated together, their circumscription has varied considerably. The third edition (1903) of Engler's ''Syllabus'' only included Campynema, but positioned it as Campynematoideae, a subfamily of Amaryllidaceae. Phylogeny The synthesis of molecular data with cladistic analysis suggests that the Liliales form one of eleven orders of monocotyledons. Sequencing of the '' rbc ...
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Colchicaceae
Colchicaceae is a family of flowering plants that includes 15 genera with a total of about 285 known species according to Christenhusz and Byng in 2016. Description The family is characterized by the presence of colchicine. Taxonomy The APG III system, of 2009 (unchanged from the APG systems, of 1998 and 2003), recognizes this family and places it in the order Liliales, in the clade monocots. It is a group of herbaceous perennials with rhizomes or corms. The Dahlgren system and the Thorne system (1992) also recognized this family, and placed it in order Liliales in superorder Lilianae in subclass Liliidae (monocotyledons) of class Magnoliopsida (angiosperms). Genera The following is an alphabetic list of genera usually included in this family : The phylogenetic classification of Vinnersten & Manning circumscribes the family as follows: * tribe Burchardieae ** ''Burchardia'' R.Br. * tribe Uvularieae ** ''Disporum'' Salisb. (excluding the North American species to the genus ''P ...
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Smilacaceae
Smilacaceae, the greenbriers, is a family of flowering plants. While they were often assigned to a more broadly defined family Liliaceae, most recent botanists have accepted the two as distinct families, diverging around 55 million years ago during the Early Paleogene. One characteristic that distinguishes Smilacaceae from most of the other members of the Liliaceae-like Liliales is that it has true vessels in its conducting tissue. Another is that the veins of the leaves, between major veins, are reticulate (net-shaped), rather than parallel as in most monocots. Taxonomy The APG II system, of 2003 (unchanged from the APG system, of 1998), recognizes this family and places it in the order Liliales, in the clade monocots. Earlier it was a family of two genera, ''Heterosmilax'' and ''Smilax'', but DNA studies have shown that ''Heterosmilax'' has arisen from ''Smilax'' and the two genera are now merged. This results in Smilax being the only genus in Smilacaceae with about 210 know ...
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APG III System
The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a further revision, the APG IV system. Along with the publication outlining the new system, there were two accompanying publications in the same issue of the Botanical Journal of the Linnean Society: * The first, by Chase & Reveal, was a formal phylogenetic classification of all land plants (embryophytes), compatible with the APG III classification. As the APG have chosen to eschew ranks above order, this paper was meant to fit the system into the existing Linnaean hierarchy for those that prefer such a classification. The result was that all land plants were placed in the class Equisetopsida, which was then divided into 16 subclasses and a multitude of superorders. * The second, by Haston ''et al.'', was a linear sequence of families followi ...
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Philesiaceae
Philesiaceae is a family of flowering plants, including two genera, each with a single species. The members of the family are woody shrubs or vines endemic to southern Chile. The APG III system, of 2009 (unchanged from the APG II system of 2003 and the APG system, of 1998) places the family in the order Liliales, in the clade monocots. They are a sister to the family Ripogonaceae which is endemic to Australia, New Zealand and New Guinea. Genera *''Lapageria'' Ruiz & Pavon *''Philesia ''Philesia'' is a South American genus of flowering plants in the family Philesiaceae first described as a genus in 1789. The flower is mostly pink in color, but some darker, more purple variations have been recorded. There is only one known spe ...'' Comm. ex Jussieu References External linksPhilesiaceaein L. Watson and M.J. Dallwitz (1992 onwards). The families of flowering plants descriptions, illustrations, identification, information retrieval.'' Version: 9 March 2006. http://delta-intk ...
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Dioscoreales
The Dioscoreales are an order of monocotyledonous flowering plants in modern classification systems, such as the Angiosperm Phylogeny Group and the Angiosperm Phylogeny Web. Within the monocots Dioscoreales are grouped in the lilioid monocots where they are in a sister group relationship with the Pandanales. The Dioscoreales must contain the family Dioscoreaceae which includes the yam ('' Dioscorea'') which is an important food source in many regions. Older systems tended to place all lilioid monocots with reticulate veined leaves (such as Smilacaceae and Stemonaceae together with Dioscoraceae) in Dioscoreales. As currently circumscribed by phylogenetic analysis using combined morphology and molecular methods, Dioscreales contains many reticulate veined vines in Dioscoraceae, it also includes the myco-heterotrophic Burmanniaceae and the autotrophic Nartheciaceae. The order consists of three families, 22 genera and about 850 species. Description Dioscoreales are vines or her ...
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Melanthiaceae
Melanthiaceae, also called the bunchflower family, is a family of flowering herbaceous perennial plants native to the Northern Hemisphere. Along with many other lilioid monocots, early authors considered members of this family to belong to the family Liliaceae, in part because both their sepals and petals closely resemble each other and are often large and showy like those of lilies, while some more recent taxonomists have placed them in a family Trilliaceae. The most authoritative modern treatment, however, the APG III system of 2009 (unchanged from the 2003 APG II system and the 1998 APG system), places the family in the order Liliales, in the clade monocots. Circumscribed in this way, the family includes up to 17 genera. Familiar members of the family include the genera ''Paris'' and ''Trillium''. Genera and species , the World Checklist of Selected Plant Families accepted 17 genera in the family.Search for "Melanthiaceae", They have been divided into five tribes. It h ...
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Ripogonaceae
''Ripogonum'' (sometimes ''Rhipogonum'') is a genus of flowering plants confined to eastern Australia, New Zealand, and New Guinea. Until recently this genus was included in the family Smilacaceae, and earlier in the family Liliaceae, but it has now been separated as its own family Ripogonaceae (sometimes Rhipogonaceae). Like most species of the closely related Smilacaceae, most species of ''Ripogonum'' are woody vines. Differences from Smilacaceae include that ''Ripogonum'' lacks stipules, it has a wet rather than dry stigma, its seeds and leaves contain starch, and its guard cells contain oil. Description The six species of ''Ripogonum'' are perennials, either vines or shrubs. The leaves, which may have several different arrangements, lack stipules. The stems may have prickles. The Australian species are bisexual; others are unisexual. Individual flowers have six white to pale green or yellow tepals. The ovary has three locules with two ovules per locule. The fruit is a ber ...
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