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Microheliella
''Microheliella'' is a monotypic genus of protists containing the sole species ''M. maris'', first described in 2012. It has a variety of unusual morphological characteristics which make its broader classification difficult. These include a centrosome with two concentric granular shells and axopodia much simpler in structure than in visually similar protists (other 'heliozoa'). Recent phylogenomic analyses suggest the microhelida are sister to the Cryptista, forming a clade called Pancryptista, which would be sister to the Archaeplastida The Archaeplastida (or kingdom Plantae ''sensu lato'' "in a broad sense"; pronounced /ɑːrkɪ'plastɪdə/) are a major group of eukaryotes, comprising the photoautotrophic red algae (Rhodophyta), green algae, land plants, and the minor group .... References {{Taxonbar, from=Q64895907 Endohelean genera ...
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Archaeplastida
The Archaeplastida (or kingdom Plantae ''sensu lato'' "in a broad sense"; pronounced /ɑːrkɪ'plastɪdə/) are a major group of eukaryotes, comprising the photoautotrophic red algae (Rhodophyta), green algae, land plants, and the minor group glaucophytes. It also includes the non-photosynthetic lineage Rhodelphidia, a predatorial (eukaryotrophic) flagellate that is sister to the Rhodophyta, and probably the microscopic picozoans. The Archaeplastida have chloroplasts that are surrounded by two membranes, suggesting that they were acquired directly through a single endosymbiosis event by feeding on a cyanobacterium. All other groups which have chloroplasts, besides the amoeboid genus ''Paulinella'', have chloroplasts surrounded by three or four membranes, suggesting they were acquired secondarily from red or green algae. Unlike red and green algae, glaucophytes have never been involved in secondary endosymbiosis events. The cells of the Archaeplastida typically lack centriol ...
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Protist
A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exclusion of other eukaryotes means that protists do not form a natural group, or clade. Therefore, some protists may be more closely related to animals, plants, or fungi than they are to other protists. However, like the groups ''algae'', ''invertebrates'', and '' protozoans'', the biological category ''protist'' is used for convenience. Others classify any unicellular eukaryotic microorganism as a protist. The study of protists is termed protistology. History The classification of a third kingdom separate from animals and plants was first proposed by John Hogg in 1860 as the kingdom Protoctista; in 1866 Ernst Haeckel also proposed a third kingdom Protista as "the kingdom of primitive forms". Originally these also included prokaryotes, b ...
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Centrosome
In cell biology, the centrosome (Latin centrum 'center' + Greek sōma 'body') (archaically cytocentre) is an organelle that serves as the main microtubule organizing center (MTOC) of the animal cell, as well as a regulator of cell-cycle progression. The centrosome provides structure for the cell. The centrosome is thought to have evolved only in the metazoan lineage of eukaryotic cells. Fungi and plants lack centrosomes and therefore use other structures to organize their microtubules. Although the centrosome has a key role in efficient mitosis in animal cells, it is not essential in certain fly and flatworm species. Centrosomes are composed of two centrioles arranged at right angles to each other, and surrounded by a dense, highly structured mass of protein termed the pericentriolar material (PCM). The PCM contains proteins responsible for microtubule nucleation and anchoring — including γ-tubulin, pericentrin and ninein. In general, each centriole of the centrosome is based ...
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Pseudopodia
A pseudopod or pseudopodium (plural: pseudopods or pseudopodia) is a temporary arm-like projection of a eukaryotic cell membrane that is emerged in the direction of movement. Filled with cytoplasm, pseudopodia primarily consist of actin filaments and may also contain microtubules and intermediate filaments. Pseudopods are used for motility and ingestion. They are often found in amoebas. Different types of pseudopodia can be classified by their distinct appearances. Lamellipodia are broad and thin. Filopodia are slender, thread-like, and are supported largely by microfilaments. Lobopodia are bulbous and amoebic. Reticulopodia are complex structures bearing individual pseudopodia which form irregular nets. Axopodia are the phagocytosis type with long, thin pseudopods supported by complex microtubule arrays enveloped with cytoplasm; they respond rapidly to physical contact. Some pseudopodial cells are able to use multiple types of pseudopodia depending on the situation: Most of t ...
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Heliozoa
Heliozoa, commonly known as sun-animalcules, are microbial eukaryotes (protists) with stiff arms ( axopodia) radiating from their spherical bodies, which are responsible for their common name. The axopodia are microtubule-supported projections from the amoeboid cell body, and are variously used for capturing food, sensation, movement, and attachment. They are similar to Radiolaria, but they are distinguished from them by lacking central capsules and other complex skeletal elements, although some produce simple scales and spines. They may be found in both freshwater and marine environments. Classification Originally the heliozoa were treated together as a formal taxon Heliozoa or Heliozoea, with the rank of class or phylum, but it has been realised that they are polyphyletic, as the various orders show notable differences and are no longer believed to be descended from a single common ancestor. Instead, "heliozoa" is regarded as a descriptive term applying to various lines of proti ...
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Phylogenomic
Phylogenomics is the intersection of the fields of evolution and genomics. The term has been used in multiple ways to refer to analysis that involves genome data and evolutionary reconstructions. It is a group of techniques within the larger fields of phylogenetics and genomics. Phylogenomics draws information by comparing entire genomes, or at least large portions of genomes. Phylogenetics compares and analyzes the sequences of single genes, or a small number of genes, as well as many other types of data. Four major areas fall under phylogenomics: * Prediction of gene function * Establishment and clarification of evolutionary relationships * Gene family evolution * Prediction and retracing lateral gene transfer. The ultimate goal of phylogenomics is to reconstruct the evolutionary history of species through their genomes. This history is usually inferred from a series of genomes by using a genome evolution model and standard statistical inference methods (e.g. Bayesian inference or ...
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Cryptista
Cryptista is a clade of algae-like eukaryotes. It is most likely related to Archaeplastida which includes plants and many algae, within the larger group Diaphoretickes. Although it has sometimes placed along with Haptista in the group Hacrobia, within the kingdom Chromista, most recent studies have found that Hacrobia is not a clade. For example, in 2016, a broad phylogenomic study found that cryptists fall within the group Archaeplastida, while haptophytes are closely related to the SAR supergroup. Taxonomy Based on studies done by Cavalier-Smith, Chao & Lewis 2015 *''Corbihelia'' ** Phylum Microheliellida Tedersoo 2017 [Endohelia Cavalier-Smith 2015] *** Class Endohelea Cavalier-Smith 2012 * Clade Cryptista s.s. ** Phylum Palpitophyta Tedersoo 2017 *** Class Palpitea Cavalier-Smith 2012 ** Clade Rollomonadia Cavalier-Smith 2013 stat. nov. *** Phylum Kathablepharidophyta Okamoto & Inouye 2005 [Leucocrypta Cavalier-Smith 2015] **** Class Leucocryptea Cavalier-Smith 2004 [Kathab ...
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Picozoa
Picozoa, Picobiliphyta, Picobiliphytes, or Biliphytes are protists of a phylum of marine unicellular heterotrophic eukaryotes with a size of less than about 3 micrometers. They were formerly treated as eukaryotic algae and the smallest member of photosynthetic picoplankton before it was discovered they don't perform photosynthesis. The first species identified therein is ''Picomonas judraskeda''. They probably belong in the Archaeplastida as sister of the Rhodophyta. They were formerly placed within the cryptomonads-haptophytes assemblage. Discovery At the end of the 1990s the European project "Picodiv" clarified which organisms occur in picoplankton. In addition, for a period of two years, samples were taken in the Atlantic, in the Mediterranean, before the coast of Scotland, Alaska and Norway. Picobiliphyta were found particularly within the nutrient-poor ranges from cold coastal seas, where they can constitute up to 50 percent of the biomass. Affinities to other organisms ...
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Rhodelphidia
''Rhodelphis'' is a single-celled archaeplastid that lives in aquatic environments and is the sister group to red algae and possibly Picozoa. While red algae have no flagellated stages and are generally photoautotrophic, ''Rhodelphis'' is a flagellated predator containing a non-photosynthetic plastid. This group is important to the understanding of plastid evolution because they provide insight into the morphology and biochemistry of early archaeplastids. ''Rhodelphis'' contains a remnant plastid that is not capable of photosynthesis, but may play a role in biochemical pathways in the cell like heme synthesis and iron-sulfur clustering. The plastid does not have a genome, but genes are targeted to it from the nucleus. ''Rhodelphis'' is ovoid with a tapered anterior end bearing two perpendicularly-oriented flagella. History ''Rhodelphis'' was described by Gawryluk ''et al.'' in 2019 along with species, ''R. marinus'' and ''R. limneticus,'' which were first collected in 2015 and 2 ...
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Rhodophyta
Red algae, or Rhodophyta (, ; ), are one of the oldest groups of eukaryotic algae. The Rhodophyta also comprises one of the largest phyla of algae, containing over 7,000 currently recognized species with taxonomic revisions ongoing. The majority of species (6,793) are found in the Florideophyceae (class), and mostly consist of multicellular, marine algae, including many notable seaweeds. Red algae are abundant in marine habitats but relatively rare in freshwaters. Approximately 5% of red algae species occur in freshwater environments, with greater concentrations found in warmer areas. Except for two coastal cave dwelling species in the asexual class Cyanidiophyceae, there are no terrestrial species, which may be due to an evolutionary bottleneck in which the last common ancestor lost about 25% of its core genes and much of its evolutionary plasticity. The red algae form a distinct group characterized by having eukaryotic cells without flagella and centrioles, chloroplasts tha ...
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Chloroplastida
Viridiplantae (literally "green plants") are a clade of eukaryotic organisms that comprise approximately 450,000–500,000 species and play important roles in both terrestrial and aquatic ecosystems. They are made up of the green algae, which are primarily aquatic, and the land plants (embryophytes), which emerged from within them. Green algae traditionally excludes the land plants, rendering them a paraphyletic group. However it is accurate to think of land plants as a kind of algae. Since the realization that the embryophytes emerged from within the green algae, some authors are starting to include them. They have cells with cellulose in their cell walls, and primary chloroplasts derived from endosymbiosis with cyanobacteria that contain chlorophylls a and b and lack phycobilins. In some classification systems, the group has been treated as a kingdom, under various names, e.g. Viridiplantae, Chlorobionta, or simply Plantae, the latter expanding the traditional plant kingdom t ...
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Glaucophyta
The glaucophytes, also known as glaucocystophytes or glaucocystids, are a small group of unicellular algae found in freshwater and moist terrestrial environments, less common today than they were during the Proterozoic. The stated number of species in the group varies from about 14 to 26. Together with the red algae (Rhodophyta) and the green algae plus land plants (Viridiplantae or Chloroplastida), they form the Archaeplastida. However, the relationships among the red algae, green algae and glaucophytes are unclear, in large part due to limited study of the glaucophytes. The glaucophytes are of interest to biologists studying the development of chloroplasts because some studies suggest they may be similar to the original algal type that led to green plants and red algae in that glaucophytes may be basal Archaeplastida. Unlike red and green algae, glaucophytes only have asexual reproduction. Characteristics The plastids of glaucophytes are known as 'muroplasts', 'cyanopla ...
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