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Mertensophryne Howelli
''Mertensophryne howelli'' (common name: Mrora forest toad) is a species of toad in the family Bufonidae. It is endemic to the coast of Tanzania and known from the Mafia Island and Zanzibar. The species is named after professor Kim Howell for his contributions to the herpetology of Tanzania. Description The holotype, an adult female, measured in snout to vent length and in snout to urostyle tip length. The male paratype measured in snout to urostyle tip length. The maximum reported female snout–vent length is and clutch size 60 eggs. The top of the head is flat, as typical for species of the formerly recognized genus ''Stephopuedes''. Parotoid glands are flattened and broad. Dorsal and lateral skin of head and parotoid region are densely covered with sharply pointed, light-tipped spines, which are especially densely packed on canthus and lateral edge of eyelids. Spines on loreal region are fewer and smaller. Colouration is generally dark brown, with a light upper lip and ...
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Toad
Toad is a common name for certain frogs, especially of the family Bufonidae, that are characterized by dry, leathery skin, short legs, and large bumps covering the parotoid glands. A distinction between frogs and toads is not made in scientific taxonomy, but is common in popular culture (folk taxonomy), in which toads are associated with drier, rougher skin and more terrestrial habitats. List of toad families In scientific taxonomy, toads include the true toads (Bufonidae) and various other terrestrial or warty-skinned frogs. Non-bufonid "toads" can be found in the families: * Bombinatoridae ( fire-bellied toads and jungle toads) * Calyptocephalellidae (helmeted water toad and false toads) * Discoglossidae ( midwife toads) * Myobatrachidae (Australian toadlets) * Pelobatidae (European spadefoot toad) * Rhinophrynidae ( burrowing toads) * Scaphiopodidae (American spadefoot toads) * Microhylidae ( narrowmouth toads) Biology Usually the largest of the bumps on the skin ...
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Bufonidae
{{Cat main, Toad This category contains both species commonly called toads, and the true toads from the family Bufonidae {{Cat main, Toad This category contains both species commonly called toads, and the true toads from the family Bufonidae {{Cat main, Toad This category Category, plural categories, may refer to: Philosophy and general uses *Categorization, cat .... Animals by common name Frogs ...
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Endemic
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can be also be referred to as an ''endemism'' or in scientific literature as an ''endemite''. For example '' Cytisus aeolicus'' is an endemite of the Italian flora. '' Adzharia renschi'' was once believed to be an endemite of the Caucasus, but it was later discovered to be a non-indigenous species from South America belonging to a different genus. The extreme opposite of an endemic species is one with a cosmopolitan distribution, having a global or widespread range. A rare alternative term for a species that is endemic is "precinctive", which applies to ...
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Tanzania
Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and the Indian Ocean to the east; Mozambique and Malawi to the south; Zambia to the southwest; and Rwanda, Burundi, and the Democratic Republic of the Congo to the west. Mount Kilimanjaro, Africa's highest mountain, is in northeastern Tanzania. According to the United Nations, Tanzania has a population of million, making it the most populous country located entirely south of the equator. Many important hominid fossils have been found in Tanzania, such as 6-million-year-old Pliocene hominid fossils. The genus Australopithecus ranged across Africa between 4 and 2 million years ago, and the oldest remains of the genus ''Homo'' are found near Lake Olduvai. Following the rise of '' Homo erectus'' 1.8 million years ago, humanity spread ...
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Mafia Island
Mafia Island (Kisiwa cha Mafia) is an island and district of Pwani Region, Tanzania. The island is the third largest in Tanzanian ocean territory, but is not administratively included within the semi-autonomous region of Zanzibar, which has been politically separate since 1890. Mafia Island forms the major part of Mafia District, one of the six administrative districts in the Pwani coastal region. The local archipelago and the main island are sometimes called Chole shamba in Swahili, hinterlands of the former maintown of Chole on Mafia Bay. The island's population is over 40,000. The economy is based on fishing, subsistence agriculture and the market in Kilindoni. The island attracts some tourists, mainly scuba divers, birdwatchers, game fishermen, and people wanting relaxation. Geography Mafia Island, (), lies off the east coast of Tanzania opposite the delta of the Rufiji River. The wide stretch of water between the delta and the island is called the Mafia Channel. Mafi ...
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Zanzibar
Zanzibar (; ; ) is an insular semi-autonomous province which united with Tanganyika in 1964 to form the United Republic of Tanzania. It is an archipelago in the Indian Ocean, off the coast of the mainland, and consists of many small islands and two large ones: Unguja (the main island, referred to informally as Zanzibar) and Pemba Island. The capital is Zanzibar City, located on the island of Unguja. Its historic centre, Stone Town, is a World Heritage Site. Zanzibar's main industries are spices, raffia and tourism. In particular, the islands produce cloves, nutmeg, cinnamon, and black pepper. For this reason, the Zanzibar Archipelago, together with Tanzania's Mafia Island, are sometimes referred to locally as the "Spice Islands". Tourism in Zanzibar is a more recent activity, driven by government promotion that caused an increase from 19,000 tourists in 1985, to 376,000 in 2016. The islands are accessible via 5 ports and the Abeid Amani Karume International Airport, w ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Paratype
In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Often there is more than one paratype. Paratypes are usually held in museum research collections. The exact meaning of the term ''paratype'' when it is used in zoology is not the same as the meaning when it is used in botany. In both cases however, this term is used in conjunction with ''holotype''. Zoology In zoological nomenclature, a paratype is officially defined as "Each specimen of a type series other than the holotype.", ''International Code of Zoological Nomenclature'' In turn, this definition relies on the definition of a "type series". A type series is the material (specimens of organisms) that was cited in the original publication of the new species or subspecies, and was not excluded from being type material by the author (th ...
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Parotoid Gland
The parotoid gland (alternatively, paratoid gland) is an external skin gland on the back, neck, and shoulder of toads and some frogs and salamanders. It can secrete a number of milky alkaloid substances (depending on the species) known collectively as bufotoxins, which act as neurotoxins to deter predation. These cutaneous glands are called parotoid as they are somewhat similarly positioned to mammalian parotid gland, although the latter have a different function, excreting saliva within the mouth rather than externally excreted defensive chemicals. A study of the parotoid glands of the Colorado River toad in 1976 found that the parotoid glands were "composed of numerous lobules", each of which is a separate unit with a lumen surrounded by a double cell layer. The cell layers have interlocking microvilli. The study found that the outer cell layer resembled smooth muscle cells, with some organelles hypothesised to "function in some aspects of venom synthesis, active cellular trans ...
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Lore (anatomy)
The lore (adj. loreal) is the region between the eyes and nostrils of birds, reptiles, and amphibians. Ornithology In ornithology, the lore is the region between the eye and bill on the side of a bird's head. This region is sometimes featherless, and the skin may be tinted, as in many species of the cormorant family. This area, which is directly in front of the eye, features a "loral stripe" in many bird species including the red-capped plover. Herpetology In amphibians and reptiles, lore pertains to the regions immediately adjacent to the eyes and between the eyes and nostrils. These are analogous to the lore on birds which corresponds to the region between the eye and the beak. In snakes and reptiles, a loreal scale also refers to the scales which lie between the eye and the nostril. In crotaline snakes (pit vipers), loreal pits are present on either side of the head.Campbell JA, Lamar WW. 2004. The Venomous Reptiles of the Western Hemisphere. 2 volumes. Comstock Publishi ...
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Jozani Chwaka Bay National Park
The Jozani Chwaka Bay National Park is a national park in Tanzania located on the island of Zanzibar. It is the only national park in Zanzibar. The Zanzibar red colobus, '' Procolobus kirkii'' (its population count is about 1000) found in the park, a rain forest species (unlike the black-and-white colobus found in other regions of Africa), is also known as Kirk's red colobus, named after Sir John Kirk, the British Resident of Zanzibar who had first brought it to the attention of zoological science. It is now adopted as the flagship species for conservation in Zanzibar, from the mid-1990s. Other species of fauna found in the park are the Sykes monkey, bush babies, more than 50 species of butterfly and 40 species of birds. The nocturnal Zanzibar tree hyrax, which has four ‘toes’ on its front feet and three on its back, is said to be the first hyrax species that has acclimatized to the forest. As part of the tourism circuit, the park attracts 10% of the over 100,000 visitors ...
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Mertensophryne
''Mertensophryne'' is a genus of true toads (family Bufonidae). They are found in eastern and southern Democratic Republic of Congo to Kenya, Tanzania, Malawi, southeastern Zimbabwe, and adjacent Mozambique. Their common names include snouted frogs, Chirinda forest toads, and forest toads. The genus is named for Robert Mertens, German zoologist and herpetologist. Taxonomy ''Mertensophryne'', as currently understood, consist of the former ''"Bufo" taitanus'' group and the genera ''Mertensophryne'' (as formerly defined) and ''Stephopaedes''. The latter is monophyletic and still recognized as a subgenus. The closest relatives of ''Mertensophryne'' remain uncertain but probably include ''Poyntonophrynus'', '' Vandijkophrynus'', and ''Capensibufo''. Description ''Mertensophryne'' lack tympanum and columella Lucius Junius Moderatus Columella (; Arabic: , 4 – ) was a prominent writer on agriculture in the Roman Empire. His ' in twelve volumes has been completely preserved a ...
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