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Machairoceratops
''Machairoceratops'' (meaning "bent sword horned face"), previously known as the "Wahweap centrosaurine B", is an extinct genus of centrosaurine ceratopsian dinosaur known from the Late Cretaceous Wahweap Formation (late Campanian stage) of Grand Staircase–Escalante National Monument, southern Utah, United States. Discovery It contains a single species, ''M. cronusi'', first described and named in 2016 by Eric K. Lund, Patrick M. O’Connor, Mark A. Loewen and Zubair A. Jinnah. The generic name is derived from Greek ''machairis'', meaning "bent sword", in reference to its unique frill ornamentation showing two forward curving horns on the frill's uppermost part, and Latinized Greek ''ceratops'', meaning "horned-face", which is a common suffix for ceratopsian genera names. The specific name ''cronusi'' refers to Cronus, a Greek god who deposed his father Uranus by castrating him with a sickle or scythe based on the mythology, and as such is shown carrying a curved bladed wea ...
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Machairoceratops NT Small
''Machairoceratops'' (meaning "bent sword horned face"), previously known as the "Wahweap centrosaurine B", is an extinct genus of centrosaurine ceratopsian dinosaur known from the Late Cretaceous Wahweap Formation (late Campanian stage) of Grand Staircase–Escalante National Monument, southern Utah, United States. Discovery It contains a single species, ''M. cronusi'', first described and named in 2016 by Eric K. Lund, Patrick M. O’Connor, Mark A. Loewen and Zubair A. Jinnah. The generic name is derived from Greek ''machairis'', meaning "bent sword", in reference to its unique frill ornamentation showing two forward curving horns on the frill's uppermost part, and Latinized Greek ''ceratops'', meaning "horned-face", which is a common suffix for ceratopsian genera names. The specific name ''cronusi'' refers to Cronus, a Greek god who deposed his father Uranus by castrating him with a sickle or scythe based on the mythology, and as such is shown carrying a curved bladed wea ...
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Machairoceratops Squamosal
''Machairoceratops'' (meaning "bent sword horned face"), previously known as the "Wahweap centrosaurine B", is an extinct genus of centrosaurine ceratopsian dinosaur known from the Late Cretaceous Wahweap Formation (late Campanian stage) of Grand Staircase–Escalante National Monument, southern Utah, United States. Discovery It contains a single species, ''M. cronusi'', first described and named in 2016 by Eric K. Lund, Patrick M. O’Connor, Mark A. Loewen and Zubair A. Jinnah. The generic name is derived from Greek ''machairis'', meaning "bent sword", in reference to its unique frill ornamentation showing two forward curving horns on the frill's uppermost part, and Latinized Greek ''ceratops'', meaning "horned-face", which is a common suffix for ceratopsian genera names. The specific name ''cronusi'' refers to Cronus, a Greek god who deposed his father Uranus by castrating him with a sickle or scythe based on the mythology, and as such is shown carrying a curved bladed wea ...
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Centrosaurinae
Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithischians. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Alberta, Montana, and Alaska) but isolated taxa have been found in China and Utah as well. Defining features of centrosaurines include a large nasal horn, short supratemporal horns, and an ornamented frill projecting from the back of the skull. With the exception of ''Centrosaurus apertus'', all adult centrosaurines have spike-like ornaments midway up the skull. Morphometric analysis shows that centrosaurines differ from other ceratopsian groups in skull, snout, and frill shapes. There is evidence to suggest that male centrosaurines had an extended period of adolescence, and sexual ornamentation did not appear until adulthood. Centrosaurinae was named by paleontologist Lawrence Lambe in 1915, with ''Centrosaurus'' as the type genus. The cen ...
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Wahweap Formation
The Wahweap Formation of the Grand Staircase–Escalante National Monument is a geological formation in southern Utah and northern Arizona, around the Lake Powell region, whose strata date back to the Late Cretaceous (Campanian stage). Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, North America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 574–588. . Age The Wahweap formation is divided into four members, which are listed below with their respective ages: * Last Chance Creek Member: 82.17-81.55 Ma * Reynolds Point Member: 81.55-80.61 Ma * Coyote Point Member: 80.61-79 Ma * Pardner Canyon Member: 79-77.29 Ma Paleobiota Invertebrates The Wahweap Formation shows a substantial amount of invertebrate activity ranging from fossilized insect burrows in petrified logs to var ...
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Centrosaurini
Centrosaurinae (from the Greek language, Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal Ornithischia, ornithischians. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Alberta, Montana, and Alaska) but isolated taxa have been found in China and Utah as well. Defining features of centrosaurines include a large nasal horn, short supratemporal horns, and an ornamented frill projecting from the back of the skull. With the exception of ''Centrosaurus, Centrosaurus apertus'', all adult centrosaurines have spike-like ornaments midway up the skull. Morphometric analysis shows that centrosaurines differ from other ceratopsian groups in skull, snout, and frill shapes. There is evidence to suggest that male centrosaurines had an extended period of adolescence, and sexual Biological ornament, ornamentation did not appear until adulthood. Centrosaurinae was named by paleontologist Lawrence ...
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Centrosaurine
Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithischians. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Alberta, Montana, and Alaska) but isolated taxa have been found in China and Utah as well. Defining features of centrosaurines include a large nasal horn, short supratemporal horns, and an ornamented frill projecting from the back of the skull. With the exception of ''Centrosaurus apertus'', all adult centrosaurines have spike-like ornaments midway up the skull. Morphometric analysis shows that centrosaurines differ from other ceratopsian groups in skull, snout, and frill shapes. There is evidence to suggest that male centrosaurines had an extended period of adolescence, and sexual ornamentation did not appear until adulthood. Centrosaurinae was named by paleontologist Lawrence Lambe in 1915, with ''Centrosaurus'' as the type genus. The cen ...
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Maximum Parsimony (phylogenetics)
In phylogenetics, maximum parsimony is an optimality criterion under which the phylogenetic tree that minimizes the total number of character-state changes (or miminizes the cost of differentially weighted character-state changes) is preferred. Under the maximum-parsimony criterion, the optimal tree will minimize the amount of homoplasy (i.e., convergent evolution, parallel evolution, and evolutionary reversals). In other words, under this criterion, the shortest possible tree that explains the data is considered best. Some of the basic ideas behind maximum parsimony were presented by James S. Farris in 1970 and Walter M. Fitch in 1971. Maximum parsimony is an intuitive and simple criterion, and it is popular for this reason. However, although it is easy to ''score'' a phylogenetic tree (by counting the number of character-state changes), there is no algorithm to quickly ''generate'' the most-parsimonious tree. Instead, the most-parsimonious tree must be sought in "tree space" ...
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Bayesian Inference In Phylogeny
Bayesian inference of phylogeny combines the information in the prior and in the data likelihood to create the so-called posterior probability of trees, which is the probability that the tree is correct given the data, the prior and the likelihood model. Bayesian inference was introduced into molecular phylogenetics in the 1990s by three independent groups: Bruce Rannala and Ziheng Yang in Berkeley, Bob Mau in Madison, and Shuying Li in University of Iowa, the last two being PhD students at the time. The approach has become very popular since the release of the MrBayes software in 2001, and is now one of the most popular methods in molecular phylogenetics. Bayesian inference of phylogeny background and bases Bayesian inference refers to a probabilistic method developed by Reverend Thomas Bayes based on Bayes' theorem. Published posthumously in 1763 it was the first expression of inverse probability and the basis of Bayesian inference. Independently, unaware of Bayes' work, Pierre- ...
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Uranus (mythology)
In Greek mythology, Uranus ( ), sometimes written Ouranos ( grc, Οὐρανός, , sky, ), is the personification of the sky and one of the Greek primordial deities. According to Hesiod, Uranus was the son and husband of Gaia (Earth), with whom he fathered the first generation of Titans. However, no cult addressed directly to Uranus survived into Classical times, and Uranus does not appear among the usual themes of Greek painted pottery. Elemental Earth, Sky, and Styx might be joined, however, in solemn invocation in Homeric epic. Uranus is associated with the Roman god Caelus and the Jewish god Yahweh. Etymology Most linguists trace the etymology of the name to a Proto-Greek form ''*Worsanós'' (), enlarged from *''ṷorsó-'' (also found in Greek ''()'' 'to urinate', Sanskrit ''varṣá'' 'rain', Hittite ''ṷarša-'' 'fog, mist').Robert S. P. Beekes, ''Etymological Dictionary of Greek'', vol. 2 (Leiden: Brill, 2009), 1128–1129. The basic Indo-European root is ''* ...
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Parietal Bone
The parietal bones () are two bones in the Human skull, skull which, when joined at a fibrous joint, form the sides and roof of the Human skull, cranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four borders, and four angles. It is named from the Latin ''paries'' (''-ietis''), wall. Surfaces External The external surface [Fig. 1] is convex, smooth, and marked near the center by an eminence, the parietal eminence (''tuber parietale''), which indicates the point where ossification commenced. Crossing the middle of the bone in an arched direction are two curved lines, the superior and inferior temporal lines; the former gives attachment to the temporal fascia, and the latter indicates the upper limit of the muscular origin of the temporal muscle. Above these lines the bone is covered by a tough layer of fibrous tissue – the epicranial aponeurosis; below them it forms part of the temporal fossa, and affords attachment to the temporal muscle. ...
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Squamosal Bone
The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestral component of the dermal roof and is typically thin compared to other skull bones. The squamosal bone lies ventral to the temporal series and otic notch, and is bordered anteriorly by the postorbital. Posteriorly, the squamosal articulates with the quadrate and pterygoid bones. The squamosal is bordered anteroventrally by the jugal and ventrally by the quadratojugal. Function in reptiles In reptiles, the quadrate and articular bones of the skull articulate to form the jaw joint. The squamosal bone lies anterior to the quadrate bone. Anatomy in synapsids Non-mammalian synapsids In non-mammalian synapsids, the jaw is composed of four bony elements and referred to as a quadro-articular jaw because the joint is between the articular an ...
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Braincase
In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skullcap. The remainder of the skull is the facial skeleton. In comparative anatomy, neurocranium is sometimes used synonymously with endocranium or chondrocranium. Structure The neurocranium is divided into two portions: * the membranous part, consisting of flat bones, which surround the brain; and * the cartilaginous part, or chondrocranium, which forms bones of the base of the skull. In humans, the neurocranium is usually considered to include the following eight bones: * 1 ethmoid bone * 1 frontal bone * 1 occipital bone * 2 parietal bones * 1 sphenoid bone * 2 temporal bones The ossicles (three on each side) are usually not included as bones of the neurocranium. There may variably also be extra sutural bones present. Below the ne ...
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