''Machairoceratops'' (meaning "bent sword horned face"), previously known as the "Wahweap centrosaurine B", is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

centrosaurine Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithischians. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Al ...

ceratopsia Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

known from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

Wahweap Formation The Wahweap Formation of the Grand Staircase–Escalante National Monument is a geological formation in southern Utah and northern Arizona, around the Lake Powell region, whose strata date back to the Late Cretaceous (Campanian stage). Dinosaur r ...

(late

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

stage) of

Grand Staircase–Escalante National Monument The Grand Staircase–Escalante National Monument (GSENM) is a United States national monument protecting the Grand Staircase, the Kaiparowits Plateau, and the Canyons of the Escalante (Escalante River) in southern Utah. It was established in 199 ...

, southern

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

Discovery

It contains a single species, ''M. cronusi'', first described and named in

2016 File:2016 Events Collage.png, From top left, clockwise: Bombed-out buildings in Ankara following the 2016 Turkish coup d'état attempt; the impeachment trial of Brazilian President Dilma Rousseff; Damaged houses during the 2016 Nagorno-Karabakh ...

by Eric K. Lund, Patrick M. O’Connor, Mark A. Loewen and Zubair A. Jinnah. The generic name is derived from

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

''machairis'', meaning "bent sword", in reference to its unique frill ornamentation showing two forward curving horns on the frill's uppermost part, and

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

ized Greek ''ceratops'', meaning "horned-face", which is a common suffix for ceratopsian genera names. The specific name ''cronusi'' refers to

Cronus In Ancient Greek religion and mythology, Cronus, Cronos, or Kronos ( or , from el, Κρόνος, ''Krónos'') was the leader and youngest of the first generation of Titans, the divine descendants of the primordial Gaia (Mother Earth) and ...

, a Greek god who deposed his father

Uranus Uranus is the seventh planet from the Sun. Its name is a reference to the Greek god of the sky, Uranus (mythology), Uranus (Caelus), who, according to Greek mythology, was the great-grandfather of Ares (Mars (mythology), Mars), grandfather ...

by castrating him with a sickle or scythe based on the mythology, and as such is shown carrying a curved bladed weapon. ''Machairoceratops'' is known solely from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

UMNH VP 20550, found in 2006, which is housed at the

Natural History Museum of Utah The Natural History Museum of Utah (NHMU) is a museum located in Salt Lake City, Utah, United States. The museum shows exhibits of natural history subjects, with an emphasis on Utah and the Intermountain West. The mission of the museum is to il ...

. It is represented by a partial

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

including two curved and elongate eyesocket horncores, the left

jugal bone The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anato ...

, a nearly complete but slightly deformed

braincase In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skul ...

, the left

squamosal bone The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestral c ...

, and a

parietal bone The parietal bones () are two bones in the Human skull, skull which, when joined at a fibrous joint, form the sides and roof of the Human skull, cranium. In humans, each bone is roughly quadrilateral in form, and has two surfaces, four borders, an ...

complex and its unique horn ornamentation, all collected in association.

Phylogeny

Lund ''et al.'' (2016) tested the position of ''Machairoceratops'' within

Centrosaurinae Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithischians. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Al ...

by performing

maximum parsimony In phylogenetics, maximum parsimony is an optimality criterion under which the phylogenetic tree that minimizes the total number of character-state changes (or miminizes the cost of differentially weighted character-state changes) is preferred. ...

and Bayesian phylogenetic species level analyses. The maximum parsimony analysis yielded a large

polytomy An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tr ...

at the base of Centrosaurinae, with only

Centrosaurini Centrosaurinae (from the Greek language, Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal Ornithischia, ornithischians. Centrosaurine fossil remains are known primarily from the northern regio ...

, most of

Pachyrhinosaurini Pachyrhinosaurini is a tribe (biology), tribe of Centrosaurinae, centrosaurine dinosaurs. The clade existed during the Late Cretaceous, about 84.9 to 68.5 million years ago, evolving during the earliest Campanian, and becoming extinct i ...

(''

Einiosaurus ''Einiosaurus'' is a genus of herbivorous centrosaurine ceratopsian dinosaur from the Upper Cretaceous (Campanian stage) of northwestern Montana. The name means 'buffalo lizard', in a combination of Blackfeet Indian ''eini'' and Latinized Ancient ...

'', ''

Wendiceratops ''Wendiceratops'' is a genus of herbivorous centrosaurine ceratopsian dinosaur from the Late Cretaceous of Canada. Discovery In 2010, Canadian fossil hunter Wendy Sloboda found a centrosaurine bonebed in the Pinhorn Provincial Grazing Reser ...

'' and

Pachyrostra Pachyrhinosaurini is a tribe of centrosaurine dinosaurs. The clade existed during the Late Cretaceous, about 84.9 to 68.5 million years ago, evolving during the earliest Campanian, and becoming extinct in the Maastrichtian. The tribe co ...

), and a clade formed by ''

Avaceratops ''Avaceratops'' is a genus of small herbivorous ceratopsian dinosaurs which lived during the late Campanian during the Late Cretaceous Period in what are now the Northwest United States. Most fossils come from the Judith River Formation. Discove ...

'' and ''

Nasutoceratops ''Nasutoceratops'' is an extinct genus of ceratopsian dinosaur. It is a basal centrosaurine which lived during the Late Cretaceous Period (late Campanian, about 76.0-75.5 Ma). Fossils have been found in southern Utah, United States. ''Nasutocer ...

'' being resolved. The Bayesian analysis yielded a fully resolved topology which is shown below. The

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

presented below follows a more recent phylogenetic analysis by Chiba ''et al.'' (2017):

References

{{Taxonbar, from=Q24091673 Centrosaurines Dinosaurs of North America Fossil taxa described in 2016 Paleontology in Utah Ornithischian genera

Discovery

Phylogeny

See also

References