Lohuecotitan
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Lohuecotitan
''Lohuecotitan'' is an extinct genus of titanosaurian sauropod dinosaur which lived during the Late Cretaceous in Spain. The only species known in the genus is ''Lohuecotitan pandafilandi'', described and named in 2016.. Discovery and naming The fossil remains of ''Lohuecotitan'' were discovered in the site of Lo Hueco, Fuentes, Cuenca, which is part of the Villalba de la Sierra Formation. The formation dates from the Upper Campanian to the Lower Maastrichtian, and would have represented a muddy coastal floodplain. Multiple partial sauropod skeletons have been discovered at this site, with the teeth and braincases recovered representing at least two distinct types of titanosaur. The holotype specimen of ''Lohuecotitan'', HUE-EC-01, is a disarticulated partial skeleton consisting of cervical, dorsal, sacral, and caudal vertebrae, ribs, an ulna, both ischia, a pubis, a femur, a fibula, and a tibia, along with some indeterminate remains. The name of ''Lohuecotitan'' combines a r ...
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Lohuecotitan Model - Castilla-La Mancha Paleontological Museum 03
''Lohuecotitan'' is an extinct genus of titanosaurian sauropod dinosaur which lived during the Late Cretaceous in Spain. The only species known in the genus is ''Lohuecotitan pandafilandi'', described and named in 2016.. Discovery and naming The fossil remains of ''Lohuecotitan'' were discovered in the site of Lo Hueco, Fuentes, Cuenca, which is part of the Villalba de la Sierra Formation. The formation dates from the Upper Campanian to the Lower Maastrichtian, and would have represented a muddy coastal floodplain. Multiple partial sauropod skeletons have been discovered at this site, with the teeth and braincases recovered representing at least two distinct types of titanosaur. The holotype specimen of ''Lohuecotitan'', HUE-EC-01, is a disarticulated partial skeleton consisting of cervical, dorsal, sacral, and caudal vertebrae, ribs, an ulna, both ischia, a pubis, a femur, a fibula, and a tibia, along with some indeterminate remains. The name of ''Lohuecotitan'' combines a r ...
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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a '' nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ...
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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Villalba De La Sierra Formation
The Villalba de la Sierra Formation is a Campanian to Maastrichtian geologic formation in Spain. Fossil dinosaur eggs have been reported from the formation, that comprises gypsiferous, grey, argillaceous mudstones and sandstones, deposited in a floodplain environmentWeishampel et al., 2004, pp.517-607 characterised by high seasonality and variability in water availability. Fossil content The formation has provided abundant titanosaurian remains, including '' Lohuecotitan'' were found in the formation.Díez Díaz et al., 2016 More than 10,000 fossil remains of various fishes, amphibians, lizards, dinosaurs ('' Ampelosaurus sp.'', '' Rhabdodon sp.''), turtles ('' Foxemys mechinorum'', '' Iberoccitanemys convenarum''), and crocodiles ('' Lohuecosuchus megadontos'', '' Agaresuchus fontisensis'', '' Musturzabalsuchus sp.'') are also known from the site, one of the richest for the Late Cretaceous in Europe.Ortega et al., 2015 See also * List of dinosaur-bearing rock formations ** L ...
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Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 (± 0.2) to 72.1 (± 0.2) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian. The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks. Etymology The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the department of Charente-Maritime. The original type locality was a series of outcrop near the village of Aubeterre-sur-Dronne in the same region. Definition The base of the Campanian Stage is defined as a place in the stratigraphic column wher ...
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Jainosaurus
''Jainosaurus'' is a genus of titanosaurian sauropod dinosaur of India and wider Asia, which lived in the Maastrichtian (approximately 68 mya (unit), million years ago). No accurate estimate of the length, height, or weight has yet been made. The humerus of the type specimen is 134 centimetres long. Etymology The specific name (zoology), specific name of ''J. septentrionalis'' means "northern" in Latin, a reference to the fact that the species was discovered on the Northern hemisphere whereas ''Antarctosaurus'' means "saurian from the Southern hemisphere" because its type species ''Antarctosaurus wichmannianus'' was found in Argentina. The generic name honours the Indian paleontologist Sohan Lal Jain, who worked on the cranial nerve impressions in the skull; and in 1982 published a study about the results. Ironically, Jain himself considered the remains synonymous with ''Titanosaurus'' in the 1997 description of ''Isisaurus''. However, Wilson and Upchurch (2003) rejected th ...
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Articular Processes
The articular processes or zygapophyses (Greek ζυγον = "yoke" (because it links two vertebrae) + απο = "away" + φυσις = "process") of a vertebra are projections of the vertebra that serve the purpose of fitting with an adjacent vertebra. The actual region of contact is called the ''articular facet''.Moore, Keith L. et al. (2010) ''Clinically Oriented Anatomy'', 6th Ed, p.442 fig. 4.2 Articular processes spring from the junctions of the pedicles and laminæ, and there are two right and left, and two superior and inferior. These stick out of an end of a vertebra to lock with a zygapophysis on the next vertebra, to make the backbone more stable. * The superior processes or prezygapophysis project upward from a lower vertebra, and their articular surfaces are directed more or less backward (oblique coronal plane). * The inferior processes or postzygapophysis project downward from a higher vertebra, and their articular surfaces are directed more or less forward and outwa ...
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Saltasaurus
''Saltasaurus'' (which means "lizard from Salta") is a genus of saltasaurid dinosaur of the Late Cretaceous period of Argentina. Small among sauropods, though still heavy by the standards of modern creatures, ''Saltasaurus'' was characterized by a short neck and stubby limbs. It was the first genus of sauropod known to possess armour of bony plates embedded in its skin. Such small bony plates, called osteoderms, have since been found on other titanosaurians. Discovery The fossils of ''Saltasaurus'' were excavated by José Bonaparte, Martín Vince and Juan C. Leal between 1975 and 1977 at the Estancia "El Brete". The find was in 1977 reported in the scientific literature. ''Saltasaurus'' was named and described by Bonaparte and Jaime E. Powell in 1980. The type species is ''Saltasaurus loricatus''. Its generic name is derived from Salta Province, the region of north-west Argentina where the first fossils were recovered. The specific name means "protected by small armoured plat ...
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Vertebral Column
The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordata, chordates has been replaced by a segmented series of bone: vertebrae separated by intervertebral discs. Individual vertebrae are named according to their region and position, and can be used as anatomical landmarks in order to guide procedures such as Lumbar puncture, lumbar punctures. The vertebral column houses the spinal canal, a cavity that encloses and protects the spinal cord. There are about 50,000 species of animals that have a vertebral column. The human vertebral column is one of the most-studied examples. Many different diseases in humans can affect the spine, with spina bifida and scoliosis being recognisable examples. The general structure of human vertebrae is fairly typical of that found in mammals, reptiles, and birds. Th ...
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Skeletal Pneumaticity
Skeletal pneumaticity is the presence of air spaces within bones. It is generally produced during development by excavation of bone by pneumatic diverticula (air sacs) from an air-filled space, such as the lungs or nasal cavity. Pneumatization is highly variable between individuals, and bones not normally pneumatized can become pneumatized in pathological development. Cranial pneumaticity Pneumatization occurs in the skulls of mammals, crocodilians and birds among extant tetrapods. Pneumatization has been documented in extinct archosaurs including dinosaurs and pterosaurs. Pneumatic spaces include the paranasal sinuses and some of the mastoid cells. Postcranial pneumaticity Postcranial pneumaticity is found largely in certain archosaur groups, namely dinosaurs, pterosaurs, and birds. Vertebral pneumatization is widespread among saurischian dinosaurs, and some theropods have quite widespread pneumatization, for example ''Aerosteon riocoloradensis'' has pneumatization of the ilium ...
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Somphospondyli
Somphospondyli is an extinct clade of titanosauriform sauropods that lived from the Late Jurassic until the end of the Late Cretaceous, comprising all titanosauriforms more closely related to Titanosauria proper than Brachiosauridae. The remains of somphospondylans have been discovered in all six continents. Classification The group has officially been defined under the PhyloCode as the largest clade containing ''Saltasaurus loricatus'', but not ''Giraffatitan brancai.'' Features found as diagnostic of this clade by Mannion ''et al.'' (2013) include the possession of at least 15 cervical vertebrae; a bevelled radius bone end; sacral vertebrae with camellate internal texture; convex posterior articular surfaces of middle to posterior caudal vertebrae; biconvex distal caudal vertebrae; humerus anterolateral corner "squared"; among multiple others. The following cladogram depicts the reference phylogeny used to defined Somphospondlyi under the PhyloCode The ''International Code ...
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