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Isoëtes
''Isoetes'', commonly known as the quillworts, is the only extant genus of plants in the family Isoetaceae, which is in the class of lycopods. There are currently 192 recognized species, with a cosmopolitan distribution but with the individual species often scarce to rare. Some botanists split the genus, separating two South American species into the genus ''Stylites'', although molecular data place these species among other species of ''Isoetes'', so that ''Stylites'' does not warrant taxonomic recognition. Species of ''Isoetes'' virtually identical to modern forms have existed since the Jurassic epoch. The name of the genus may also be spelled ''Isoëtes''. The diaeresis (two dots over the e) indicates that the o and the e are to be pronounced in two distinct syllables. Including this in print is optional; either spelling (''Isoetes'' or ''Isoëtes'') is correct. Description Quillworts are mostly aquatic or semi-aquatic in clear ponds and slow-moving streams, though several ( ...
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Isoetes Megasporangia L
''Isoetes'', commonly known as the quillworts, is the only extant genus of plants in the family Isoetaceae, which is in the class of lycopods. There are currently 192 recognized species, with a cosmopolitan distribution but with the individual species often scarce to rare. Some botanists split the genus, separating two South American species into the genus ''Stylites'', although molecular data place these species among other species of ''Isoetes'', so that ''Stylites'' does not warrant taxonomic recognition. Species of ''Isoetes'' virtually identical to modern forms have existed since the Jurassic epoch. The name of the genus may also be spelled ''Isoëtes''. The diaeresis (two dots over the e) indicates that the o and the e are to be pronounced in two distinct syllables. Including this in print is optional; either spelling (''Isoetes'' or ''Isoëtes'') is correct. Description Quillworts are mostly aquatic or semi-aquatic in clear ponds and slow-moving streams, though several ( ...
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Isoetes Butleri
''Isoetes butleri'', commonly known as limestone quillwort, is a species of plant in the quillwort family, a member of the lycophyte The lycophytes, when broadly circumscribed, are a vascular plant (tracheophyte) subgroup of the kingdom Plantae. They are sometimes placed in a division Lycopodiophyta or Lycophyta or in a subdivision Lycopodiophytina. They are one of the oldes ...s. It is native to the eastern United States, where it is scattered and local. It is found almost exclusively on seasonally wet alkaline soils. Common habitats include rocky prairies and glades, most often over limestone. This species is inconspicuous and easily overlooked, due to its superficial similarity to a vegetative grass or sedge. The leaves of ''Isoetes butleri'' die back at the end of spring, when its habitat becomes hot and dry. It is then reduced to a below ground corm. References {{Taxonbar, from=Q17023023 butleri Flora of the Southeastern United States Flora without expected TNC ...
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Homosporous
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into a ne ...
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Spore
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into a new s ...
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Lycopodiaceae
The Lycopodiaceae (class Lycopodiopsida, order Lycopodiales) are an old family of vascular plants, including all of the core clubmosses and firmosses, comprising 16 accepted genera and about 400 known species. This family originated about 380 million years ago in the early Devonian, though the diversity within the family has been much more recent. "Wolf foot" is another common name for this family due to the resemblance of either the roots or branch tips to a wolf's paw. Description Members of Lycopodiaceae are not spermatophytes and so do not produce seeds. Instead they produce spores, which are oily and flammable, and are the most economically important aspects of these plants. The spores are of one size (i.e. the plants are isosporous) and are borne on a specialized structure at the apex of a shoot called a strobilus (plural: strobili), which resembles a tiny battle club, from which the common name derives. Members of the family share the common feature of having a microphyll, ...
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Heterospory
Heterospory is the production of spores of two different sizes and sexes by the sporophytes of land plants. The smaller of these, the microspore, is male and the larger megaspore is female. Heterospory evolved during the Devonian period from isospory independently in several plant groups: the clubmosses, the ferns including the arborescent horsetails, and progymnosperms. This occurred as part of the process of evolution of the timing of sex differentiation.Sussex, I.M. (1966) The origin and development of heterospory in vascular plants. Chapter 9 in ''Trends in Plant morphogenesis'', ed. by E.G. Cutter, Longmans. Origin of heterospory Heterospory evolved due to natural selection that favoured an increase in propagule size compared with the smaller spores of homosporous plants. Heterosporous plants, similar to anisosporic plants, produce two different sized spores in separate sporangia that develop into separate male and female gametophytes. It is proposed that the emergence ...
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Selaginellaceae
''Selaginella'' is the sole genus of vascular plants in the family Selaginellaceae, the spikemosses or lesser clubmosses. This family is distinguished from Lycopodiaceae The Lycopodiaceae (class Lycopodiopsida, order Lycopodiales) are an old family of vascular plants, including all of the core clubmosses and firmosses, comprising 16 accepted genera and about 400 known species. This family originated about 380 mil ... (the clubmosses) by having scale-leaves bearing a ligule and by having heterospory, spores of two types. They are sometimes included in an informal paraphyletic group called the "fern ally, fern allies". ''Selaginella moellendorffii, S. moellendorffii'' is an important model organism. Its genome has been sequenced by the United States Department of Energy's Joint Genome Institute. The name ''Selaginella'' was erected by Palisot de Beauvois solely for the species ''Selaginella selaginoides'', which turns out (with the closely related ''Selaginella deflexa'') to ...
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Lepidodendron
''Lepidodendron'' is an extinct genus of primitive vascular plants belonging to the family Lepidodendraceae, part of a group of Lycopodiopsida known as scale trees or arborescent lycophytes, related to Isoetes, quillworts and Lycopodiopsida, lycopsids (club mosses). They were part of the coal forest flora. They sometimes reached heights of , and the trunks were often over in diameter. They thrived during the Carboniferous Period (358.9 to 298.9 million years ago). Sometimes erroneously called "giant club mosses", the genus was actually more closely related to modern quillworts than to modern club mosses. Within the form classification system used within paleobotany, ''Lepidodendron'' is both used for the whole plant as well as specifically the stems and leaves. Etymology The name ''Lepidodendron'' comes from the Greek language, Greek wikt:λεπίς, λεπίς ', scale, and wikt:δένδρον, δένδρον ''dendron'', tree. Growth During the early stages of growth, ''L ...
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Angiosperms
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils are in the ...
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Chromosomes
A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are the histones. These proteins, aided by chaperone proteins, bind to and condense the DNA molecule to maintain its integrity. These chromosomes display a complex three-dimensional structure, which plays a significant role in transcriptional regulation. Chromosomes are normally visible under a light microscope only during the metaphase of cell division (where all chromosomes are aligned in the center of the cell in their condensed form). Before this happens, each chromosome is duplicated (S phase), and both copies are joined by a centromere, resulting either in an X-shaped structure (pictured above), if the centromere is located equatorially, or a two-arm structure, if the centromere is located distally. The joined copies are now called sis ...
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Non-coding DNA
Non-coding DNA (ncDNA) sequences are components of an organism's DNA that do not encode protein sequences. Some non-coding DNA is transcribed into functional non-coding RNA molecules (e.g. transfer RNA, microRNA, piRNA, ribosomal RNA, and regulatory RNAs). Other functional regions of the non-coding DNA fraction include regulatory sequences that control gene expression; scaffold attachment regions; origins of DNA replication; centromeres; and telomeres. Some non-coding regions appear to be mostly nonfunctional such as introns, pseudogenes, intergenic DNA, and fragments of transposons and viruses. Fraction of non-coding genomic DNA In bacteria, the coding regions typically take up 88 % of the genome. The remaining 12 % consists largely of non-coding genes and regulatory sequences, which means that almost all of the bacterial genome has a function. The amount of coding DNA in eukaryrotes is usually a much smaller fraction of the genome because eukaryotic genomes contai ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is homoplasy. The recurrent evolution of flight is a classic example, as flying insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas '' homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different traits. Convergent evolution is similar to parallel evo ...
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