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Holospira
''Holospira'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod molluscs in the family Urocoptidae. ''Holospira'' is the type genus of the subfamily 'Holospirinae Pilsbry, 1946'. Etymology ''Holos'' (ὅλος) from Ancient Greek means "‘whole’" or "‘entire’" and ''spira'' (spīra) from Latin means spire. Shell description The shell is small, cylindric, terminating above in a conic spire, retaining all the whorls, rimate or perforate. The shell has 11-21 whorls, which are closely coiled. The first 1½ of whorls are smooth. The rest of whorls are smooth, striate or ribbed. The suture is superficial. Body whorl is more or less built forward. The aperture is small, obliquely pear-shaped, rounded or oval. The peristome is expanded or reflected, continuous and usually free throughout. The columella (internal column) is hollow, variously sculptured or smooth. Anatomy The foot is small, narrow for its length. The lung is long and narrow ...
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Holospira Elizabethae Columella
''Holospira'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod molluscs in the family Urocoptidae. ''Holospira'' is the type genus of the subfamily 'Holospirinae Pilsbry, 1946'. Etymology ''Holos'' (ὅλος) from Ancient Greek means "‘whole’" or "‘entire’" and ''spira'' (spīra) from Latin means spire. Shell description The shell is small, cylindric, terminating above in a conic spire, retaining all the whorls, rimate or perforate. The shell has 11-21 whorls, which are closely coiled. The first 1½ of whorls are smooth. The rest of whorls are smooth, striate or ribbed. The suture is superficial. Body whorl is more or less built forward. The aperture is small, obliquely pear-shaped, rounded or oval. The peristome is expanded or reflected, continuous and usually free throughout. The columella (internal column) is hollow, variously sculptured or smooth. Anatomy The foot is small, narrow for its length. The lung is long an ...
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Holospira Elizabethae
''Holospira elizabethae'' is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusc in the family Urocoptidae. Paratypes of this species are in the collection of the Natuurhistorisch Museum Rotterdam.NMR Collection database
. accessed 19 June 2009


Original description

''Holospira elizabethae'' was originally discovered and described by in 1889. The type locality is Amula village, which is between the towns of and

Urocoptidae
Urocoptidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea. Taxonomy 2005 taxonomy The family Urocoptidae was classified in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005): * Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868 * Apomatinae Paul, 1982 * Brachypodellinae H. B. Baker, 1956 * Eucalodiinae P. Fischer & Crosse, 1873 * Holospirinae Pilsbry, 1946 * Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972 * Tetrentodontinae Bartsch, 1943 2008 taxonomy Uit de Weerd (2008) moved the Urocoptidae to the newly established superfamily Urocoptoidea based on molecular phylogeny research. The following cladogram based on Bayesian analy ...
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Urocoptoidea
Urocoptoidea is a Superfamily (zoology), superfamily of land snails, gastropods in the suborder Helicina (suborder), Helicina. MolluscaBase eds. (2020). MolluscaBase. Urocoptoidea Pilsbry, 1898 (1868). Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=932607 on 2020-08-25 Families In 2005, according to the Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), taxonomy of the Gastropoda by Bouchet & Rocroi, the families Urocoptidae and Cerionidae were classified within the superfamily Orthalicoidea . In 2008, the superfamily Urocoptoidea was established by the malacologist Uit de Weerd, based on molecular phylogeny research.Uit de Weerd D. R. (2008). "Delimitation and phylogenetics of the highly diverse land snail family Urocoptidae (Gastropoda, Pulmonata) based on 28S rRNA sequence data: A reunion with ''Cerion''". ''Journal of Molluscan Studies'' 74: 317-329. . In 2012, ThompsonThompson F. G. (2012). "The land snail g ...
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Pulmonate
Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includes many land and freshwater families, and several marine families. The taxon Pulmonata as traditionally defined was found to be polyphyletic in a molecular study per Jörger ''et al.'', dating from 2010. Pulmonata are known from the Carboniferous Period to the present. Pulmonates have a single atrium and kidney, and a concentrated, symmetrical, nervous system. The mantle cavity is located on the right side of the body, and lacks gills, instead being converted into a vascularised lung. Most species have a shell, but no operculum, although the group does also include several shell-less slugs. Pulmonates are hermaphroditic, and some groups possess love darts. Linnean taxonomy The taxonomy of this group according to the taxonomy of the Ga ...
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Body Whorl
The body whorl is part of the morphology of the shell in those gastropod mollusks that possess a coiled shell. The term is also sometimes used in a similar way to describe the shell of a cephalopod mollusk. In gastropods In gastropods, the body whorl, or last whorl, is the most recently formed and largest whorl (or revolution) of a spiral or helical shell, terminating in the aperture. It is called the "body whorl" because most of the body of the soft parts of the animal fits into this whorl. The proportional size of the body whorl in gastropod shells differs greatly according to the actual shell morphology. For shells in which the rate of whorl expansion of each revolution around the axis is very high, the aperture and the body whorl are large, and the shell tends to be low spired. The shell of the abalone is a good example of this kind of shell. The opposite tendency can sometimes create a high spire with very little whorl increase per revolution. In these instances, e.g. ...
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Suture (gastropod)
In anatomy, a suture is a fairly rigid joint between two or more hard elements of an organism, with or without significant overlap of the elements. Sutures are found in the skeletons or exoskeletons of a wide range of animals, in both invertebrates and vertebrates. Sutures are found in animals with hard parts from the Cambrian period to the present day. Sutures were and are formed by several different methods, and they exist between hard parts that are made from several different materials. Vertebrate skeletons The skeletons of vertebrate animals (fish, amphibians, reptiles, birds, and mammals) are made of bone, in which the main rigid ingredient is calcium phosphate. Cranial sutures The skulls of most vertebrates consist of sets of bony plates held together by cranial sutures. These sutures are held together mainly by Sharpey's fibers which grow from each bone into the adjoining one. Sutures in the ankles of land vertebrates In the type of crurotarsal ankle which is found i ...
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Whorl (mollusc)
A whorl is a single, complete 360° revolution or turn in the spiral growth of a mollusc shell. A spiral configuration of the shell is found in numerous gastropods, but it is also found in shelled cephalopods including ''Nautilus'', ''Spirula'' and the large extinct subclass of cephalopods known as the ammonites. A spiral shell can be visualized as consisting of a long conical tube, the growth of which is coiled into an overall helical or planispiral shape, for reasons of both strength and compactness. The number of whorls which exist in an adult shell of a particular species depends on mathematical factors in the geometric growth, as described in D'Arcy Wentworth Thompson's classic 1917 book ''On Growth and Form'', and by David Raup. The main factor is how rapidly the conical tube expands (or flares-out) over time. When the rate of expansion is low, such that each subsequent whorl is not that much wider than the previous one, then the adult shell has numerous whorls. When the ...
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Gastropod Shell
The gastropod shell is part of the body of a Gastropoda, gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less (slugs) but may have a remnant within the mantle, or in some cases the shell is reduced such that the body cannot be retracted within it (semi-slug). Some snails also possess an operculum that seals the opening of the shell, known as the Aperture (mollusc), aperture, which provides further protection. The study of mollusc shells is known as conchology. The biological study of gastropods, and other molluscs in general, is malacology. Shell morphology terms vary by species group. Shell layers The gastropod shell has three major layers secreted by the Mantle (mollusc), mantle. The calcareous central layer, tracum, is typically made of calcium carbonate precipitated into an organic matrix known as c ...
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Spire (mollusc)
A spire is a part of the coiled shell of molluscs. The spire consists of all of the whorls except for the body whorl. Each spire whorl represents a rotation of 360°. A spire is part of the shell of a snail, a gastropod mollusc, a gastropod shell, and also the whorls of the shell in ammonites, which are fossil shelled cephalopods. In textbook illustrations of gastropod shells, the tradition (with a few exceptions) is to show most shells with the spire uppermost on the page. The spire, when it is not damaged or eroded, includes the protoconch (also called the nuclear whorls or the larval shell), and most of the subsequent teleoconch whorls (also called the postnuclear whorls), which gradually increase in area as they are formed. Thus the spire in most gastropods is pointed, the tip being known as the "apex". The word "spire" is used, in an analogy to a church spire or rock spire, a high, thin, pinnacle. The "spire angle" is the angle, as seen from the apex, at which a spire ...
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Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the Roman Republic it became the dominant language in the Italian region and subsequently throughout the Roman Empire. Even after the fall of Western Rome, Latin remained the common language of international communication, science, scholarship and academia in Europe until well into the 18th century, when other regional vernaculars (including its own descendants, the Romance languages) supplanted it in common academic and political usage, and it eventually became a dead language in the modern linguistic definition. Latin is a highly inflected language, with three distinct genders (masculine, feminine, and neuter), six or seven noun cases (nominative, accusative, genitive, dative, ablative, and vocative), five declensions, four verb conjuga ...
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Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic period (), and the Classical period (). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Epic and Classical periods of the language. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regarded as a separate historical stage, although its earliest form closely resembles Attic Greek and its latest form approaches Medieval Greek. There were several regional dialects of Ancient Greek, of which Attic Greek developed into Koine. Dia ...
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