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Urocoptidae is a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of air-breathing land snails,
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate gastropod
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the superfamily Urocoptoidea.


Taxonomy


2005 taxonomy

The family Urocoptidae was classified in the superfamily
Orthalicoidea The Orthalicoidea are a superfamily of air-breathing land snails, terrestrial gastropod mollusks in the infraorder Orthalicoidei of the suborder HelicinaMolluscaBase eds. (2021). MolluscaBase. Simpulopsidae Schileyko, 1999. Accessed through: W ...
(according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005): * Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868 * Apomatinae Paul, 1982 * Brachypodellinae H. B. Baker, 1956 * Eucalodiinae P. Fischer & Crosse, 1873 * Holospirinae Pilsbry, 1946 * Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972 * Tetrentodontinae Bartsch, 1943


2008 taxonomy

Uit de Weerd (2008) moved the Urocoptidae to the newly established superfamily Urocoptoidea based on
molecular phylogeny Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
research. The following
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
based on
Bayesian analysis Bayesian inference is a method of statistical inference in which Bayes' theorem is used to update the probability for a hypothesis as more evidence or information becomes available. Bayesian inference is an important technique in statistics, and e ...
of sequences of
28S ribosomal RNA 28S ribosomal RNA is the structural ribosomal RNA (rRNA) for the large subunit (LSU) of eukaryotic cytoplasmic ribosomes, and thus one of the basic components of all eukaryotic cells. It has a size of 25S in plants and 28S in mammals, hence th ...
shows the phylogenic relations within the Urocoptoidea. There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.


2012 taxonomy

Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails ('' Epirobia'', '' Propilsbrya'', '' Pectinistemma'', '' Gyrocion'', '' Prionoloplax''), that were previously placed within Urocoptidae.Thompson F. G. (2012). "The land snail genus ''Epirobia'' and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". '' Bulletin Florida Museum of Natural History'' 51: 167-215
PDF
Data from
molecular phylogeny Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.


Genera

Genera in the family Urocoptidae include: Urocoptinae * '' Autocoptis'' Pilsbry, 1902 * ''
Brachypodella ''Brachypodella'' is a genus of gastropods belonging to the family Urocoptidae. The species of this genus are found in Northern and Central America. Species Species: *''Brachypodella antiperversa'' *''Brachypodella bourguignatiana'' *''Bra ...
'' L. Pfeiffer, 1840 (as ''Cylindrella'') * '' Urocoptis'' Beck, 1837 - type genus of the family Urocoptidae Apominae * '' Apoma'' Beck, 1837 - type genus of the subfamily Apomatinae * '' Simplicervix'' Pilsbry, 1903 Brachypodellinae * ''
Brachypodella ''Brachypodella'' is a genus of gastropods belonging to the family Urocoptidae. The species of this genus are found in Northern and Central America. Species Species: *''Brachypodella antiperversa'' *''Brachypodella bourguignatiana'' *''Bra ...
'' Beck, 1837 - type genus of the subfamily Brachypodellinae Eucalodiinae * '' Anisospira'' Strebel, 1880 * '' Archegocoptis'' Pilsbry, 1903 *†'' Condonella'' McLellan, 1927 * '' Dissotropis'' Bartsch, 1906 * '' Eucalodium'' Crosse & P. Fischer, 1868 - type genus of the subfamily Eucalodiinae Holospirinae * '' Coelostemma'' Dall, 1895 * '' Hendersoniella'' Dall, 1905 * '' Holospira'' Martens, 1860 - type genus of the subfamily Holospirinae Microceraminae * '' Microceramus'' Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae * '' Macroceramus'' Guilding, 1828 Tetrentodontinae * '' Tetrentodon'' Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae * '' Torrecoptis'' Bartsch, 1943 Other genera include:http://pick4.pick.uga.edu/mp/20q?search=Urocoptidae * '' Allocoptis'' Thompson & Franz, 1976 * '' Amphicosmia'' Pilsbry & Vanatta, 1898 * '' Anoma'' * '' Arangia'' * '' Bactrocoptis'' * '' Badiofax'' * '' Bostrichocentrum'' * '' Bostrychocentrum'' * '' Callonia'' * ''
Centralia Centralia may refer to: Places Australia *Central Australia, sometimes called "Centralia" Canada * Centralia, Ontario ** RCAF Station Centralia, a former Royal Canadian Air Force training base ** Centralia (Essery Field) Aerodrome United State ...
'' * '' Cochlodinella'' * '' Coelocentrum'' * '' Geoscala'' * '' Gongylostoma'' * '' Gyraxis'' * '' Haplocion'' * '' Heterocoptis'' * '' Idiostemma'' * '' Liocallonia'' * '' Metastoma'' * '' Mychostoma'' * '' Nesocoptis'' * '' Nodulia'' * '' Oligostylus'' * '' Organocoptis'' * '' Pfeiffericoptis'' * '' Pineria'' * '' Pleurostemma'' * '' Poeycoptis'' * '' Pseudopineria'' * '' Pycnoptychia'' Pilsbry & Vanatta, 1898 * '' Sagracoptis'' * '' Spiroceramus'' * '' Spirostemma'' * '' Strophina'' * '' Tomelasmus'' * '' Trilamellaxis'' * '' Uncinicoptis''


References


Further reading

* ; 7 figs. * Carlos de la Torre. 1912. ''New Cuban Urocoptis of the U. cinerea group''. The Nautilus
volume 26
number 4, page
55
58
plate 6
* Kabat A. R., Herschler R. & González G. A. (2012). "Resolution of taxonomic problems associated with the complex publication history of the seminal Torre and Bartsch monograph on Cuban Urocoptidae (Gastropoda, Pulmonata)". ''
Zootaxa ''Zootaxa'' is a peer-reviewed scientific mega journal for animal taxonomists. It is published by Magnolia Press (Auckland, New Zealand). The journal was established by Zhi-Qiang Zhang in 2001 and new issues are published multiple times a week ...
'' 3362: 43-53
PDF
* Pilsbry H. A. & Henderson J. B. jr. (February 1913). ''Two new Cuban Urocoptidae''
The Nautilus, volume 26
number 10, page
109
110. * Miguel L. Jaume & Alfredo de la Torre. 1976. ''Los Urocoptidae de Cuba (Mollusca-Pulmonata)''. Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp. * * Richardson C. L. 1991. ''Urocoptidae: catalogue of species''. Tryonia 22: 1-245. * Tryon G. W. (1903) '' Manual of Conchology. Second series: Pulmonata
Volume 15
Urocoptidae''. Continued by H. A. Pilsbry.


External links







{{Taxonbar, from=Q3074668