Hoeloceras
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Hoeloceras
''Hoeloceras'' is an extinct orthoconic nautiloid cephalopod from the upper Ordovician, generally included in the Actinocerida. Nautiloids are a subclass of shelled cephalopods that were once diverse and numerous but are now represented by only a couple of genera, '' Nautilus'' and '' Allonautilus''. ''Hoeloceras'', which has a straight shell with a broadly curved to almost flattened venter on the underside and a more highly arched dorsum on the upper, meeting acutely on either side, was named by Walter Sweet in 1958 (Chen & Liu 1977, Teichert 1964) who first recognized it in Ordovician sediments near Oslo, Norway, and included it in the Actinocerida. Teichert (1964) identified ''Hoeloceras'' as a synonym of ''Lambeoceras'', as the acutely angled dorso-ventral seam suggests. Flower (1968) referred to ''Hoeloceras'' as a "flatfish", in common with ''Lambeoceras'' and ''Gonioceras'', but reassigned it to the Reudemannoceratidae (Discosorida) noting that neither (siphuncular) an ...
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List Of Nautiloids
This list of nautiloids is a comprehensive listing of all genera that have ever been included in the subclass Nautiloidea, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (''nomina dubia''), or were not formally published (''nomina nuda''), as well as junior synonyms of more established names, and genera that are no longer considered nautiloids. Most of the listed genera are found in Part K of the ''Treatise on Invertebrate Paleontology''. Some, added since the year of publication (1964) are found simply in various scientific journals and special publications. The named genera are based on type specimens which are housed in various museums and other academic institutions worldwide, available to interested researchers. Note that ''Allonautilus'' and ''Nautilus'' are the only extant genera. A *†'' Acanthonautilus'' *†'' Acaroceras'' *†''Acleistoceras'' *†'' Acrosphaerorthoceras'' ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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Animalia
Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a bilaterally symmetric body plan. The Bilateria include the protostomes, containing animals such as nematodes, arthropods, flatworms, annelids and molluscs, and the deuterostomes, containing the echinode ...
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Mollusca
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropod ...
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Cephalopoda
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been identified. Tw ...
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Nautiloidea
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rela ...
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Actinocerida
The Actinocerida are an order of generally straight, medium to large cephalopods that lived during the early and middle Paleozoic, distinguished by a siphuncle composed of expanded segments that extend into the adjacent chambers, in which deposits formed within contain a system of radial canals and a narrow space along the inner side of the connecting ring known as a paraspatium. (Teichert 1964) Septal necks are generally short and cyrtochoanitic, some being recumbent, some hook shaped. Most grew to lengths of about but some, like the Huroniidae of the Silurian grew significantly larger. Ecology The Actinocerida inhabited shallow to quite deep waters, where they alternated between swimming and lying on the bottom. They were predatory, and able to control their buoyancy to a greater degree than their contemporaries. Derivation The derivation of the Actinocerida remains enigmatic. They first appear late in the Early Ordovician (Cassinian Stage, late Floian) with the Georginida ...
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Nautilus
The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina. It comprises six living species in two genera, the type of which is the genus ''Nautilus''. Though it more specifically refers to species ''Nautilus pompilius'', the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between 4 and 10 inches. Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle.Kümmel, B. 1964. Nautiloidae-Nautilida, in the Treatise on Invertebrate Paleontology, Geological Society of America and Univ of Kansas Press, Teichert and Moore eds. Having survived relatively u ...
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Allonautilus
The genus ''Allonautilus'' contains two species of nautiluses, which have a significantly different morphology from those placed in the sister taxon ''Nautilus''. ''Allonautilus'' is now thought to be a descendant of ''Nautilus'', rendering the latter genus paraphyletic. Live individuals of the genus have only been collected in Papua New Guinea and the Solomon Islands. Little is known about their biology because they live in deep waters, whereas the better-understood genus ''Nautilus'' lives closer to the surface. The entire family Nautilidae, including all species in the genus ''Nautilus'' and ''Allonautilus'', was listed on Appendix II of the Convention on International Trade in Endangered Species of Wild Fauna and Flora CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ... (CITE ...
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Lambeoceras
''Lambeoceras'' is a genus of large actinocerids with a convexly lenticular cross section from the Upper Ordovician of North America and the sole representative of the family Lambeoceratidae. Morphological description ''Lambeoceras'' is of medium to moderately large size with a long, straight, depressed shell, broad in cross section with the dorsum and venter both about equally convex, meeting acutely along the sides. Chambers are short, and septa are closely spaced, forming broad lobes on the upper and lower sides, which meet in sharp saddles along the sides. The siphuncle is submarginal, near the ventral side and relatively narrow. Septal necks are extremely long, brims short and recumbent. Segments are broadly expanded, connecting rings thin. Radial canals within the siphuncle from broad arcs that may bifurcate close to the parispatium. Derivation and phylogeny ''Lambeoceras'' is derived from the same stock in '' Actinoceras'' that produced ''Kochoceras'' according to Flower ( ...
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Gonioceras
''Gonioceras'' is an extinct genus of actinocerid nautiloidean cephalopods typified by a broad, low shell; flattened ventrally, convexly rounded dorsally; top and bottom meeting at an acute angle along the sides. In most the shell is rather thin, especially along the lateral portion. The aperture is contracted. Sutures from broad ventral and dorsal lobes, more narrowly rounded ventro-lateral and dorso-lateral saddles, and sharp pointed lateral lobes; more complex than in later ''Lambeoceras''. The siphuncle is typically subcentral but may be closer to the venter; armenocerid in form with short segments and very short brims and containing a straight endosiphuncular canal system. ''Gonioceras'' is derived from ''Armenoceras'' and is the type genus of Gonioceratidae. Fossils are found in marine strata of the Chazy and Blackriveran of the Middle Ordovician in the Champlain region of eastern North America and at the Paquette Rapids of the Ottawa River in Ontario. The type species ...
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Discosorida
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck. Evolution The origin of the Discosorida is unknown, thought at one timeRousseau H. Flower. 1964. The Nautiloid Order Ellesmerocerida (Cephalopods); relevant pages. Memoir 12, Ne ...
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