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Discosorida
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck. Evolution The origin of the Discosorida is unknown, thought at one timeRousseau H. Flower. 1964. The Nautiloid Order Ellesmerocerida (Cephalopods); relevant pages. Memoir 12 ...
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Discosorida
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck. Evolution The origin of the Discosorida is unknown, thought at one timeRousseau H. Flower. 1964. The Nautiloid Order Ellesmerocerida (Cephalopods); relevant pages. Memoir 12 ...
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Discosoridae
Discosoridae comprise a family of endogastric discosorids, (Cephalopoda, Nautiloidea), with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian. Shell morphology The shell in the Discosoridae is conical, expanding variably with respect to genus. Curvature tends to be slight with the ventral or siphuncular side slightly concave in profile (the endogastric condition), and the opposite dorsal side arched or convex in profile (another endogastric condition). The siphuncle is composed of broadly rounded segments the increase rapidly in size toward the aperture, with growth. Expansion is commonly greater than that of the shell itself, and lies ventral of the center. Septal necks are short, brims long and recumbent. Connecting rings are thin and poorly known. Bullettes at the apical end of the connecting rings, which grasp the periphery of the previous septal foramina and connect to the inside of the previous septal necks, are never swollen. Endocones a ...
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Lowoceratidae
The Lowoceratidae is a small family of discosorids, early nautiloid cephalopods, from the Middle Silurian in which the characteristic bullette is found only in early growth stages. Lowoceratids were first found in Southampton Island in the Canadian arctic. The Lowoceratidae consists so far of only two genera, ''Lowoceras'' and ''Tuylocras'', named by Foerste and Savage in 1927 and combined by Flower in 1940 as the Lowoceratidae. Both are strongly compressed, exogastric brevicones with the ventral side more narrowly rounded than the dorsal. Shells enlarge moderately to the middle of the mature body chamber, then contract gently toward the aperture. The siphuncle in each is slightly removed from the ventral margin. The siphuncle of ''Tuyloceras'', genotype ''T. percurvatum'', starts off with slender, subquadrate segments with thick rings and swollen bullettes, but later in life the segments become expanded, rings thinner, and bullettes simple and unswollen. ''Lowoceras'', gen ...
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Westonoceratidae
Westonoceratidae are exogastric, mostly compressed, Discosorida of moderate size from the Middle Ordovician to the Lower Silurian. The siphuncle is typically close to the convexly curved outer margin of the phragmocone – the chambered part of the shell – taken to be ventral but may be more central in some genera. Connecting rings are thin to moderately thick with inflated bullettes grasping the previous septal foremina. Parietal deposits within the siphuncle from endocones in advanced genera and cameral deposits are found in some. The Westonoceratidae are derived from the Cyrtogomphoceratidae through the cyrtogomphoceratid, '' Ulrichoceras''. In turn, the Westonoceratidae gave rise to a small group of discosorids, the Lowoceratidae, which form an intermediary step before the Discosoridae. Two principal lineages have been identified, both stemming from the genus '' Teichertoceras'', a derivative of ''Ulrichoceras''. One goes simply from '' Westonoceras'' to '' Winnipeg ...
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Phragmoceratidae
The Phragmoceratidae is a family of extinct nautiloid cephalopods from the Order Discosorida that lived during the latter part of the Silurian. Diagnosis Phragmoceratids are characterized by generally compressed, upwardly curved endogastric shells with slit-like apertures for egress and ventral siphuncles with broadly expanded segments, thick connecting rings, and small to vestigial bullettes. Phragmoceratids are morphologically similar in regards to their constricted apertures to the more cylindrical and orthoconic Mandaloceratidae, also discosorids, and to the oncocerid Hemiphragmoceratidae Hemiphragmoceratidae is a family of endogastrically brevconic oncocerids characterized by elaborately visored apertures in which the hyponomic sinus in mature specimens is on a spout-like process and there may be lateral and dorsal salients. (S ... (Sweet 1964); both which also come from the Silurian. Derivation The Phragmoceratidae are derived from the Ordovician Cyrtogomphocerat ...
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Mandaloceratidae
Mandaloceratidae is a family in the nautiloid cephalopod order Discosorida, from the Middle and Upper(?) Silurian characterized by short, essentially straight shells referred to as breviconic, typically with a faintly exogastric shape produced by the profile of the body chamber. Apertures vary from round to T-shaped, with a long narrow, ventral, hyponomic sinus for the maneuvering funnel. Siphuncles are commonly central or subcentral with generally broad, expanded, segments and usually thin connecting rings. The narrow hyponomic sinus, which is an opening for the water-jet funnel by which the animal could move, much as with modern squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ... and octopods, points toward the lower or ventral side. The cross bar of the "T" or the round ...
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Reudemannoceratidae
The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center. Derivation The Reudemannoceratidae first appeared at the beginning of the Middle Ordovician, North American Whiterock stage, (since replaced by the ICS Dapingian), and are restricted to the lower part of that series. (the middle Ordovician). Their origin is unknown. The siphuncles in early members contain features in the early growth stages reminiscent of the siphuncular bulbs found the archaic Plectronoceratae of the Late Cambrian. (Flower and Teichert 1957) but so far no unambiguous Lower Ordovician intermediaries have been found. Characters Reudemannoceratids are characterized by having short septal necks in the juvenile portion of the siphuncle, toward the apex of the shell, which later in life grow ...
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Cyrtogomphoceratidae
The Cyrtogomphoceratidae are a family in the cephalopod order Discosorida that comprises genera commonly with compressed, endogastrically curved shells. Siphuncles lie close to the ventral side, segments are broadly inflated, connecting rings thick and apically expanded thick bullettes. Chambers are short, separated by shallow, dish shaped septa. Apertures are generally simple. The Cyrtogomphoceratidae are derived from the discosorid family Reudemannoceratidae, probably from '' Reudemannoceras'', through the ancestral genus '' Ulrichoceras'', and have a range from the Middle Ordovician to the Lower Silurian. The familyTeicher, C. 1964. Nautiloidea-Discosorida. Treatise on Invertebrate Paleontology, Part K. Teichert and Moore, eds. includes: :*'' Cyrtogomphoceras'' :*'' Kiaeroceras'' :*'' Konglungenoceras'' :*'' Landeroceras'' :*'' Parryoceras'' :*'' Strandoceras'' :*'' Ulrichoceras'' ''Ulrichoceras'' is also considered the source for the exogastric Westonoceratidae. T ...
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Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ident ...
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Cephalopod Orders
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been identified. Tw ...
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Ellesmerocerida
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician. Morphology The Ellesmerocerida are characterized by shells that are typically small, some even tiny, with close-spaced septa and relatively large ventral siphuncles. In some genera (e.g. '' Paleoceras''), the septa are uniformly spaced. Shells of ellesmerocerids are typically smooth and compressed and vary in form. They may be breviconic (short) or longiconic (elongate), straight (orthoconic) or curved (cyrtoconic). Cyrtoconic forms are usually endogastric, with longitudinally convex ventral margins. The apeces of straight forms typically have an endogastric curvature. Some may have grown to as much as 15 cm. Siphuncle segments are tubular or concave. Septal necks are short. Connecting rings which may appear layered are thick and typically wedge shaped with their maximum width at or near where ...
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Plectronoceratida
Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.Curt Teichert, 1988. Main Features of Cephalopod Evolution. The Mollusca Vol. 12 Paleontology and Neontology of Cephalopds; Academic Pres Inc. Occurrence Plectronoceratids are known from the Upper Cambrian (upper Franconian – middle Trempealeauan) of China and North America (Minnesota, Wisconsin). Two families are recognized (Flower, 1964),Flower, Rousseau H. 1964. The Nautiloid Order Ellesmeroceratida (Cephalopoda); Memoir 12, New Mexico Bureau of Mines and Mineral Resources, Socorro, New Mexico the generally straight to endogastric Plectronoceratidae and the slightly exogastric Balkoceratidae. Diagnostic characters Members of the Plectronocerida are characterized as follows. Shells are generally small, some even tiny, laterally compressed, curved (cyrtochonic) or straight (orthoconic). Most cyrtoconic forms are endogastric, with the ventral side longitudinally concave, or the ...
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