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Heterotardigrade
The class Heterotardigrada includes tardigrades (water bears) that have cephalic appendages and legs with four separate but similar digits or claws on each. 444 species have been described. Anatomy The anatomy of the reproductive system is an important defining feature in distinguishing the different groups of tardigrades. Heterotardigrades have gonoducts that open to the outside through a preanal gonopore, rather than opening into the rectum as in the only other confirmed class of tardigrades, the Eutardigrada Eutardigrada are a class of tardigrades (Tardigrada) without lateral appendices. Primarily freshwater bound, some species have secondarily gained the ability to live in marine environments (''Halobiotus''). By cryptobiosis many species are able t .... The third class, Mesotardigrada, is represented by a single species known from a single specimen that is now lost, and the location from which that specimen was collected has since been destroyed by an earthquake, so it ...
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Heterotardigrada
The class Heterotardigrada includes tardigrades (water bears) that have cephalic appendages and legs with four separate but similar digits or claws on each. 444 species have been described. Anatomy The anatomy of the reproductive system is an important defining feature in distinguishing the different groups of tardigrades. Heterotardigrades have gonoducts that open to the outside through a preanal gonopore, rather than opening into the rectum as in the only other confirmed class of tardigrades, the Eutardigrada. The third class, Mesotardigrada Mesotardigrada is one of three classes of tardigrades, consisting of a single species, ''Thermozodium esakii''. The animal reportedly has six claws of equal length at each foot. This species was described in 1937 by German zoologist Gilbert Rah ..., is represented by a single species known from a single specimen that is now lost, and the location from which that specimen was collected has since been destroyed by an earthquake, so its ...
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Phase-contrast Microscopy
__NOTOC__ Phase-contrast microscopy (PCM) is an optical microscopy technique that converts phase shifts in light passing through a transparent specimen to brightness changes in the image. Phase shifts themselves are invisible, but become visible when shown as brightness variations. When light waves travel through a medium other than a vacuum, interaction with the medium causes the wave amplitude and phase to change in a manner dependent on properties of the medium. Changes in amplitude (brightness) arise from the scattering and absorption of light, which is often wavelength-dependent and may give rise to colors. Photographic equipment and the human eye are only sensitive to amplitude variations. Without special arrangements, phase changes are therefore invisible. Yet, phase changes often convey important information. Phase-contrast microscopy is particularly important in biology. It reveals many cellular structures that are invisible with a bright-field microscope, as exemplif ...
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Echiniscoidea
Echiniscoidea is an order of tardigrades, a phylum of water-dwelling, eight-legged, segmented micro-animals. It was first described by Richters in 1926.Echiniscoidea
World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive list of names of marine organisms. Content The content of the registry is edited and maintained by scientific specialist ...


Families

The order Echiniscoidea consists of the following families:


References



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Order (biology)
Order ( la, wikt:ordo#Latin, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. Fo ...
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Mesotardigrada
Mesotardigrada is one of three classes of tardigrades, consisting of a single species, ''Thermozodium esakii''. The animal reportedly has six claws of equal length at each foot. This species was described in 1937 by German zoologist Gilbert Rahm from a hot spring near Nagasaki, Japan. The inability of taxonomists to replicate Rahm's finding has cast doubt on the accuracy of the description, making ''T. esakii'', and by extension the entire class Mesotardigrada, a ''taxon inquirendum''. Taxonomic ambiguity The type specimen Rahm used as the basis of his description has either been lost or it was never preserved in the first place, which Grothman ''et al''. (2017) suggest is consistent with the lax taxonomic standards of the 1930s. Thus, re-examination of the original specimen is not possible. Complicating matters further, the type locality from which Rahm collected his specimen may have been destroyed by an earthquake and subsequent searches for additional specimens matching ...
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Eutardigrada
Eutardigrada are a class of tardigrades (Tardigrada) without lateral appendices. Primarily freshwater bound, some species have secondarily gained the ability to live in marine environments (''Halobiotus''). By cryptobiosis many species are able to live temporarily in very dry environments. More than 700 species have been described. The order Apochela consists of only one family, Milnesiidae, with two genera: ''Milnesium'' and ''Limmenius''. ''Milnesium tardigradum'' can be found worldwide and is one of the biggest species among tardigrades (up to 1.4 mm); similar-looking species have been found in Cretaceous amber. The mouth of this predator has a wide opening, so the animal can eat rotifers and larger protist A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exc ...s. Other eutardi ...
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Tardigrade
Tardigrades (), known colloquially as water bears or moss piglets, are a phylum of eight-legged segmented micro-animals. They were first described by the German zoologist Johann August Ephraim Goeze in 1773, who called them Kleiner Wasserbär ("little water bear"). In 1777, the Italian biologist Lazzaro Spallanzani named them Tardigrada (), which means "slow steppers". They have been found in diverse regions of Earth's biospheremountaintops, the deep sea, tropical rainforests, and the Antarctic. Tardigrades are among the most resilient animals known, with individual species able to survive extreme conditions – such as exposure to extreme temperatures, extreme pressures (both high and low), air deprivation, radiation, dehydration, and starvation – that would quickly kill most other known forms of life. Tardigrades have survived exposure to outer space. There are about 1,300 known species in the phylum Tardigrada, a part of the superphylum Ecdysozoa consisting of animals th ...
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Class (biology)
In biological classification, class ( la, classis) is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are life, domain, kingdom, phylum, order, family, genus, and species, with class fitting between phylum and order. History The class as a distinct rank of biological classification having its own distinctive name (and not just called a ''top-level genus'' ''(genus summum)'') was first introduced by the French botanist Joseph Pitton de Tournefort in his classification of plants that appeared in his ''Eléments de botanique'', 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of construction, ...
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Oreellidae
''Oreella'' is a genus of tardigrade Tardigrades (), known colloquially as water bears or moss piglets, are a phylum of eight-legged segmented micro-animals. They were first described by the German zoologist Johann August Ephraim Goeze in 1773, who called them Kleiner Wasserbär ...s in the family Oreellidae. It is the only genus of that family. It was named and described by James Murray in 1920; the family Oreellidae was named and described by Giuseppe Ramazzotti in 1962. Ramazzotti gave the first modern description of the genus with his findings on ''O. minor'', which was differentiated from another species by the name of ''O. mollis'' by its smaller size, spur presence on internal claws, and briefer stylets. Characteristics such as these can be considered as insufficient evidence of the delineation between species, however the differences in size of ''O. minor'' (≤170µm) and ''O. mollis'' (≤251µm), as well as the presence of the aforementioned claw spurs, would be ...
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Echiniscoididae
The Echiniscoididae are a family of tardigrades, water-dwelling, eight-legged, segmented micro-animals. It is one of the four families in the Echiniscoidea order.Echiniscoididae
World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive list of names of marine organisms. Content The content of the registry is edited and maintained by scientific specialist ...
The family of Echiniscoididae consists of the following subfamilies and genera:
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Echiniscidae
The Echiniscidae are a family of tardigrades, a phylum of water-dwelling, eight-legged, segmented micro-animals. It is one of the four families in the order Echiniscoidea.Echiniscidae
The family was named by Gustav Thulin in 1928.Thulin, ''Über die Phylogenie und das System der Tardigraden'' (''On the Phylogeny and the System of Tardigrade''). Hereditas (1928) vol. 11, no. 2/3, p. 207-266


Genera

The family Echiniscidae consists of the following genera: * ''
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Carphaniidae
''Carphania fluviatilis'' is a species of tardigrade. It is the only known species within the genus '' Carphania'', which is the only genus in the family Carphaniidae, which is part of the order Echiniscoidea. It was first described by Maria Grazia Binda in 1978. It is a freshwater species, endemic in Italy.''Carphania fluviatilis'' Binda, 1978
World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive list of names of marine organisms. Content The content of the registry is edited and maintained by scientific specialist ...
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