Gnathosaurinae
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Gnathosaurinae
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "char ...
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Plataleorhynchus
''Plataleorhynchus'' is a genus of ctenochasmatid pterodactyloid pterosaur from the Late Jurassic to Early Cretaceous periods ( Tithonian to Berriasian stages) of what is now the Purbeck Limestone of Dorset, England. History and etymology The genus was named in 1995 by Stafford Howse and Andrew Milner. The type species is ''Plataleorhynchus streptophorodon''. The genus name is derived from ''Platalea'', the spoonbill, and Greek , "snout", in reference to the distinctive form of the front of the skull. The specific name is derived from Greek , "collared", and , "tooth", referring to the tooth form. ''Plataleorhynchus'' is based on holotype NHML R.11957 (earlier BMNH R.11957), an incomplete anterior upper jaw with teeth found in a chalkstone quarry near Langton Matravers. The fossil is present on a plate; its underside is visible. This jaw is notable because it expands to form a circular, spatula-like shape at the front, holding 22 narrow teeth that point sideways. Forty oth ...
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Huanhepterus
''Huanhepterus'' is an extinct genus of ctenochasmatid pterodactyloid pterosaur from the Early Cretaceous period of what is now Qingyang, Gansu, China. History The genus was named by Dong Zhiming in 1982. The type species is ''Huanhepterus quingyangensis''. The genus name refers to the Huang Jian (not the Yellow River or "Huang He", but a smaller tributary of the Jinghe River in Gansu), and combines it with a Latinized Greek ''pteron'', "wing". The specific name refers to Qinyang County. ''Huanhepterus'' is based on the holotype specimen, IVPP V9070, a partial articulated skeleton consisting mostly of impressions of the left half of the body and the beak-end of the skull. The fossil was in May 1978 found in a quarry operated by the Sanshilipu-commune, when an explosion exposed a vertebra. Its force obliterated the right half of the specimen. IVPP V9070 hails from the Early Cretaceous-age Huachihuanhe Formation of the Zhidan Group. Description ''Huanhepterus'' had a long, ...
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Gnathosaurus
''Gnathosaurus'' (meaning "jawed lizard") is a genus of ctenochasmatid pterosaur containing two species: ''G. subulatus'', named in 1833 from the Solnhofen Limestone of Germany, and ''G. macrurus'', known from the Purbeck Limestone of the UK. Its fossil remains dated back to the Late Jurassic period. History of discovery Fragments of ''Gnathosaurus'' jaws were first discovered in 1832 in the Solnhofen limestones of southern Germany but were mistaken for a piece of teleosaurid crocodile jaw by Georg zu Münster, who first named the species ''Crocodilus multidens'' in that year. Soon afterwards, Hermann von Meyer classified the same specimen as the new genus and species ''Gnathosaurus subulatus'', a name which came to be universally used shortly thereafter. In the 1860s, scientists such as Albert Oppel compared the ''G. subulatus'' jaw fragment to contemporary pterosaurs such as ''Pterodactylus'' and ''Ctenochasma'', and concluded that it was also probably a "flying reptile" rathe ...
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Eosipterus
''Eosipterus'' is an extinct genus of ctenochasmatid pterosaur from the Early Cretaceous period of Liaoning, China. Fossil remains of ''Eosipterus'' dated back to the Aptian stage of the Early Cretaceous, 124.6 million years ago. History and etymology The genus was named in 1997 by Ji Shu'an and Ji Qiang. The type species is ''Eosipterus yangi''. The genus name is derived from Greek ''eos'', "dawn" and Greek ''pteron'', "wing" with a Latin ending; and a grammar error: normally the combination would have resulted in "eopterus". The "dawn" element refers to its age but also to China being "in the east". The specific name honours paleontologist Yang Daihuan. The genus is based on the holotype specimen, GMV2117, found near Jinggangshan in western Liaoning Province, in the Yixian Formation. It was the first pterosaur discovered in that region. It consists of a partial crushed skeleton of a subadult individual on a slab, lacking skull and neck. Most vertebrae have been severely dama ...
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Feilongus
''Feilongus'' is an extinct genus of ctenochasmatid pterodactyloid pterosaur from the Barremian–Aptian-age Lower Cretaceous Yixian Formation of Beipiao, Liaoning, China. Discovery and naming The genus was named and described in 2005 by Wang Xiaolin, Alexander Kellner, Zhou Zhonghe and Diogenes de Almeida Campos. The type species is ''Feilongus youngi''. The genus name is derived from ''Feilong'', the "flying dragon". The specific name honors the Chinese paleontologist Yang Zhongjian (C. C. Young). ''Feilongus'' is based on holotype IVPP V-12539, a skull and articulated mandible, with on the same plate the detached posterior braincase, of a subadult individual. The fossil is strongly crushed. In 2014, a second specimen, DNMHM D3068 found at Gonggao, was referred to a ''Feilongus'' sp. It consists of a skull with lower jaws and four neck vertebrae. It was a possible subadult or, despite a smaller size, adult. Description The wingspan of ''Feilongus'' was estimated by Wang ...
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Pterodaustro
''Pterodaustro'' is a genus of ctenochasmatid pterodactyloid pterosaur from South America. Its fossil remains dated back to the Early Cretaceous period, about 105 million years ago. The most distinctive characteristic that separates ''Pterodaustro'' from other ctenochasmatids is its bristle-like teeth, a feature not seen in any other pterosaur. Discovery and naming The first fossils, among them the holotype PVL 2571, a thigh bone, were in the late sixties discovered by Bonaparte in the Lagarcito Formation, situated in the San Luis Province of Argentina, and dating from the Albian. The genus has later also been found in Chile in the Santa Ana Formation. At the Argentine site, the just large "Loma del ''Pterodaustro''", since then, during several expeditions, over 750 ''Pterodaustro'' specimens have been collected, 288 of them having been catalogued until 2008. This makes the species one of the best known pterosaurs, with examples from all growth stages, from egg to adult. Th ...
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Gegepterus
''Gegepterus'' is a genus of ctenochasmatid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of Liaoning, China. Only one species is known, ''G. changi''. History and etymology The genus was named in 2007 by Wang Xiaolin, Alexander Kellner, Zhou Zhonge and Diogenes de Almeida Campos. The type species is ''Gegepterus changi''. The generic name is derived from Manchu ''ge ge'', the title of a princess, in reference to the dainty gracility of the specimen, and a Latinized Greek ''pteron'', "wing". The specific name honors female paleontologist Chang Meemann, who over the years established a cordial relationship between the Chinese and Brazilian authors. In 2008 Wang emended the epithet to ''changae'', but such changes are no longer allowed by the ICZN. It is known from two specimens. The first is holotype IVPP V 11981, which was in 2001 found in grey shales from the lower part of the formation (estimated at 125 million years old), near the city of B ...
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Moganopterus
''Moganopterus'' is an extinct genus of ctenochasmatid pterosaur from the Early Cretaceous of western Liaoning Province, China. Discovery and naming The fossil of ''Moganopterus'' was discovered at the village of Xiaosanjiazi near the town of Lamadong in Liaoning Province. In 2012 it was named and described by Lü Junchang, Pu Hanyong, Xu Li, Wu Yanhua and Wei Xuefang as the type species ''Moganopterus zhuiana''. The generic name is derived from the legendary sword couple Gan Jiang and Mo Ye, in reference to the blade-like jaws, and a Latinized Greek πτερόν, ''pteron'', "wing". The specific name honors Ms. Zhu Haifen, who made the specimen available to science. The holotype, 41HIII0419, was uncovered in a layer of the Yixian Formation, dating from the Aptian, about 125 million years old. It consists of an almost complete skull with lower jaws and the second to fourth neck vertebrae. The fossil is compressed on a slab and counterslab, the splitting of the two plates havi ...
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Beipiaopterus
''Beipiaopterus'' is a genus of ctenochasmatid pterosaur from the Early Cretaceous period (Aptian stage) of what is now the Yixian Formation of the China. The genus was named in 2003 by Lü Junchang. The generic name is derived from Beipiao City in Liaoning Province and a Latinized Greek ''pteron'', "wing". The specific epithet honors paleontologist Professor Chen Peiji. The type and only species is based on holotype BPM 0002, a crushed partial skeleton of a subadult individual on a slab, missing the skull. It includes four cervical, fourteen dorsal, three sacral and nine caudal vertebrae, a complete left wing and two hind limbs. Remains of the soft parts have been preserved, including partial wing membranes, a membrane attached to the tibia, a "mane" on the neck and webbing of the feet. It had a wingspan of , and the animal itself had a length of about , however, this is only if the skull had the same length as the remainder of the body: for the neck, ten centimetres for the r ...
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Ctenochasma
''Ctenochasma'' (meaning "comb jaw") is a genus of Late Jurassic ctenochasmatid pterosaur belonging to the suborder Pterodactyloidea. Three species are currently recognized: ''C. roemeri'' (named after Friedrich Adolph Roemer), ''C. taqueti'', and ''C. elegans''. Their fossilized remains have been found in the Solnhofen Limestone of Bavaria, Germany, the "Purbeck Group" of northeastern Germany, and the Calcaires tâchetés of eastern France. History The name ''Ctenochasma'' was coined by the German paleontologist Christian Erich Hermann von Meyer in 1852, based on a single lower jaw full of closely packed teeth which he gave the species name ''Ctenochasma roemeri''.von Meyer, C.E.H. (1852). "''Ctenochasma Roemeri''." ''Paläontographica'', 2: 82–84 & pl. 13. A second species, ''C. gracile'', was named by Oppel in 1862 based on a fragmentary skull.Oppel A. (1862). "Über Fährten im lithographischen Schiefer. Paläontologische Mitteilungen aus dem Museum des Koenigl." Bayris ...
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Liaodactylus
''Liaodactylus'' is a genus of filter-feeding ctenochasmatid pterosaur from the Jurassic of China. The genus contains one species, ''L. primus'', described by Zhou ''et al.'' in 2017. As an adaptation to filter-feeding, ''Liaodactylus'' had approximately 150 long, comb-like teeth packed closely together. It is both the earliest known ctenochasmatid and the first filter-feeding pterosaur from the Jurassic Tiaojishan Formation. Later and more specialized ctenochasmatids differ from ''Liaodactylus'' in having longer snouts, smaller openings (or fenestrae) in the skull, and more teeth. Within the Ctenochasmatidae, ''Liaodactylus'' was most closely related to the European ''Ctenochasma''. Discovery and naming There is one specimen of ''Liaodactylus'' known, namely the holotype PMOL-AP00031, which is stored at the Palaeontological Museum of Liaoning. It consists of a complete skull and lower jaws, along with the first two cervical vertebrae. It originates from outcrops located about w ...
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Kepodactylus
''Kepodactylus'' is an extinct genus of ctenochasmatid pterodactyloid pterosaur from the Kimmeridgian- Tithonian-age Upper Jurassic Morrison Formation of Colorado, United States. In 1992, a team from the Denver Museum of Natural History dug up a specimen of the dinosaur ''Stegosaurus stenops'' in Garden Park, Colorado. In the quarry they also found smaller disarticulated bones from other animals, among which were those of a pterosaur new to science. In 1996, Jerald Harris and Kenneth Carpenter named the new genus. The type species is ''Kepodactylus insperatus''. The genus name is derived from Greek, ''kepos'', "garden", a reference to Garden Park and ''daktylos'', "finger", referring to the typical wing finger of pterosaurs. The specific name means "unhoped-for" in Latin, alluding to the fact that the researchers hoped to find a dinosaur, and did not expect a pterosaur. The genus is based on the holotype DMNH 21684, consisting of a cervical vertebra, humerus, several fi ...
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