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Liaodactylus
''Liaodactylus'' is a genus of filter-feeding ctenochasmatid pterosaur from the Jurassic of China. The genus contains one species, ''L. primus'', described by Zhou ''et al.'' in 2017. As an adaptation to filter-feeding, ''Liaodactylus'' had approximately 150 long, comb-like teeth packed closely together. It is both the earliest known ctenochasmatid and the first filter-feeding pterosaur from the Jurassic Tiaojishan Formation. Later and more specialized ctenochasmatids differ from ''Liaodactylus'' in having longer snouts, smaller openings (or fenestrae) in the skull, and more teeth. Within the Ctenochasmatidae, ''Liaodactylus'' was most closely related to the European ''Ctenochasma''. Discovery and naming There is one specimen of ''Liaodactylus'' known, namely the holotype PMOL-AP00031, which is stored at the Palaeontological Museum of Liaoning. It consists of a complete skull and lower jaws, along with the first two cervical vertebrae. It originates from outcrops located about w ...
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Ctenochasmatidae
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "cha ...
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Ctenochasmatid
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "ch ...
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Tiaojishan Formation
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period (Bathonian- Oxfordian stages). It is known for its exceptionally preserved fossils, including those of plants, insects and vertebrates. It is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks. Previously, the Tiaojishan Formation was grouped together with the underlying Haifanggou Formation (also known as the Jiulongshan Formation) as a single "Lanqi Formation." The Tiaojishan Formation forms a key part of the Yanliao Biota assemblage, alongside the Haifanggou Formation. Age Using Argon–argon dating, Wang and colleagues in 2005 dated part of the Tiaojishan Formation to about 160 million years ago, the beginning of the Oxfordian stage, the first stage of the Upper Jurassic epoch. In 2006, a study by Liu and colleagues used U-Pb zircon dating to conclude that the Tiaojishan Formation correlates wi ...
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Gegepterus
''Gegepterus'' is a genus of ctenochasmatid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of Liaoning, China. Only one species is known, ''G. changi''. History and etymology The genus was named in 2007 by Wang Xiaolin, Alexander Kellner, Zhou Zhonge and Diogenes de Almeida Campos. The type species is ''Gegepterus changi''. The generic name is derived from Manchu ''ge ge'', the title of a princess, in reference to the dainty gracility of the specimen, and a Latinized Greek ''pteron'', "wing". The specific name honors female paleontologist Chang Meemann, who over the years established a cordial relationship between the Chinese and Brazilian authors. In 2008 Wang emended the epithet to ''changae'', but such changes are no longer allowed by the ICZN. It is known from two specimens. The first is holotype IVPP V 11981, which was in 2001 found in grey shales from the lower part of the formation (estimated at 125 million years old), near the city of B ...
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Mandibular Symphysis
In human anatomy, the facial skeleton of the skull the external surface of the mandible is marked in the median line by a faint ridge, indicating the mandibular symphysis (Latin: ''symphysis menti'') or line of junction where the two lateral halves of the mandible typically fuse at an early period of life (1-2 years). It is not a true symphysis as there is no cartilage between the two sides of the mandible. This ridge divides below and encloses a triangular eminence, the mental protuberance, the base of which is depressed in the center but raised on either side to form the mental tubercle. The lowest (most inferior) end of the mandibular symphysis — the point of the chin — is called the "menton". It serves as the origin for the geniohyoid and the genioglossus muscles. Other animals Solitary mammalian carnivores that rely on a powerful canine bite to subdue their prey have a strong mandibular symphysis, while pack hunters delivering shallow bites have a weaker one. When filter ...
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Gnathosaurus
''Gnathosaurus'' (meaning "jawed lizard") is a genus of ctenochasmatid pterosaur containing two species: ''G. subulatus'', named in 1833 from the Solnhofen Limestone of Germany, and ''G. macrurus'', known from the Purbeck Limestone of the UK. Its fossil remains dated back to the Late Jurassic period. History of discovery Fragments of ''Gnathosaurus'' jaws were first discovered in 1832 in the Solnhofen limestones of southern Germany but were mistaken for a piece of teleosaurid crocodile jaw by Georg zu Münster, who first named the species ''Crocodilus multidens'' in that year. Soon afterwards, Hermann von Meyer classified the same specimen as the new genus and species ''Gnathosaurus subulatus'', a name which came to be universally used shortly thereafter. In the 1860s, scientists such as Albert Oppel compared the ''G. subulatus'' jaw fragment to contemporary pterosaurs such as ''Pterodactylus'' and ''Ctenochasma'', and concluded that it was also probably a "flying reptile" rathe ...
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Mandible
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movable bone of the skull (discounting the ossicles of the middle ear). It is connected to the temporal bones by the temporomandibular joints. The bone is formed prenatal development, in the fetus from a fusion of the left and right mandibular prominences, and the point where these sides join, the mandibular symphysis, is still visible as a faint ridge in the midline. Like other symphyses in the body, this is a midline articulation where the bones are joined by fibrocartilage, but this articulation fuses together in early childhood.Illustrated Anatomy of the Head and Neck, Fehrenbach and Herring, Elsevier, 2012, p. 59 The word "mandible" derives from the Latin word ''mandibula'', "jawbone" (literally "one used for chewing"), from ''wikt:mandere ...
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Angular Bone
The angular is a large bone in the lower jaw (mandible) of amphibians and reptiles (birds included), which is connected to all other lower jaw bones: the dentary (which is the entire lower jaw in mammals), the splenial, the suprangular, and the articular. It is homologous to the tympanic bone in mammals, due to the incorporation of several jaw bones into the mammalian middle ear early in mammal evolution. In therapsids (mammal ancestors and their kin), the lower jaw is made up of the dentary (the mandible in mammals) and a group of smaller "postdentary" bones near the jaw joint. As the dentary increased in size over million of years, two of these postdentary bones, the articular and angular, became increasingly reduced and the dentary eventually made direct contact with the upper jaw. These postdentary bones, even before their articular function was lost, probably transmitted sound vibrations to the stapes and, in some therapsids, a bent plate that might have supported a membrane ...
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Oxfordian (stage)
The Oxfordian is, in the ICS' geologic timescale, the earliest age of the Late Jurassic Epoch, or the lowest stage of the Upper Jurassic Series. It spans the time between 163.5 ± 1.0 Ma and 157.3 ± 1.0 Ma (million years ago). The Oxfordian is preceded by the Callovian and is followed by the Kimmeridgian. Stratigraphic definitions The Oxfordian Stage was called "Clunch Clay and Shale" by William Smith (1815–1816); in 1818 W. Buckland described them under the unwieldy title "Oxford, Forest or Fen Clay". The term Oxfordian was introduced by Alcide d'Orbigny in 1844. The name is derived from the English city of Oxford, where the beds are well developed, but they crop out almost continuously from Dorset to the coast of Yorkshire, generally forming low, broad valleys. They are well exposed at Weymouth, Oxford, Bedford, Peterborough, and in the cliffs at Scarborough, Red Cliff and Gristhorpe Bay. Rocks of this age are found also in Uig and Skye. The base of the Oxfordian Stage ...
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Jugal Bone
The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy The jugal bone is located on either side of the skull in the circumorbital region. It is the origin of several masticatory muscles in the skull. The jugal and lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones. During development, the jugal bone originates from dermal bone. In dinosaurs This bone is considered key in the determination of general traits in cases in which the entire skull has not been found intact (for instance, as with dinosaurs in paleontology). In some dinosaur genera the jugal also forms part of the lower margin of either the antorbital fenestra or the infratemporal fenestra, or both. Most commonly, this bone articu ...
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Pterodaustro
''Pterodaustro'' is a genus of ctenochasmatid pterodactyloid pterosaur from South America. Its fossil remains dated back to the Early Cretaceous period, about 105 million years ago. The most distinctive characteristic that separates ''Pterodaustro'' from other ctenochasmatids is its bristle-like teeth, a feature not seen in any other pterosaur. Discovery and naming The first fossils, among them the holotype PVL 2571, a thigh bone, were in the late sixties discovered by Bonaparte in the Lagarcito Formation, situated in the San Luis Province of Argentina, and dating from the Albian. The genus has later also been found in Chile in the Santa Ana Formation. At the Argentine site, the just large "Loma del ''Pterodaustro''", since then, during several expeditions, over 750 ''Pterodaustro'' specimens have been collected, 288 of them having been catalogued until 2008. This makes the species one of the best known pterosaurs, with examples from all growth stages, from egg to adult. Th ...
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Quadrate Bone
The quadrate bone is a skull bone in most tetrapods, including amphibians, sauropsids (reptiles, birds), and early synapsids. In most tetrapods, the quadrate bone connects to the quadratojugal and squamosal bones in the skull, and forms upper part of the jaw joint. The lower jaw articulates at the articular bone, located at the rear end of the lower jaw. The quadrate bone forms the lower jaw articulation in all classes except mammals. Evolutionarily, it is derived from the hindmost part of the primitive cartilaginous upper jaw. Function in reptiles In certain extinct reptiles, the variation and stability of the morphology of the quadrate bone has helped paleontologists in the species-level taxonomy and identification of mosasaur squamates and spinosaurine dinosaurs. In some lizards and dinosaurs, the quadrate is articulated at both ends and movable. In snakes, the quadrate bone has become elongated and very mobile, and contributes greatly to their ability to swallow very ...
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