''Kepodactylus'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

ctenochasmatid Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early ...

pterodactyloid Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...

from the

Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxford ...

- Tithonian-age Upper Jurassic Morrison Formation of

Colorado Colorado (, other variants) is a state in the Mountain states, Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the wes ...

, United States. In 1992, a team from the Denver Museum of Natural History dug up a specimen of the

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fou ...

stenops'' in Garden Park,

. In the quarry they also found smaller disarticulated bones from other animals, among which were those of a pterosaur new to science. In 1996, Jerald Harris and

Kenneth Carpenter Kenneth Carpenter (born September 21, 1949, in Tokyo, Japan) is a paleontologist. He is the former director of the USU Eastern Prehistoric Museum and author or co-author of books on dinosaurs and Mesozoic life. His main research interests ...

named the new genus. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...

is ''Kepodactylus insperatus''. The genus name is derived from Greek, ''kepos'', "garden", a reference to Garden Park and ''daktylos'', "finger", referring to the typical wing finger of pterosaurs. The specific name means "unhoped-for" in

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

, alluding to the fact that the researchers hoped to find a dinosaur, and did not expect a pterosaur. The genus is based on the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...

DMNH 21684, consisting of a

cervical In anatomy, cervical is an adjective that has two meanings: # of or pertaining to any neck. # of or pertaining to the female cervix: i.e., the ''neck'' of the uterus. *Commonly used medical phrases involving the neck are **cervical collar **cerv ...

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...

, humerus, several finger bones, and a

metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the me ...

. ''Kepodactylus'' was similar to ''

Mesadactylus ''Mesadactylus'' ('mesa finger') is an extinct genus of pterosaur from the Kimmeridgian-Tithonian-age Upper Jurassic Morrison Formation of Colorado, United States. The genus was named in 1989 by James Jensen and Kevin Padian. The type species ...

'' but larger (wingspan around 2.5 m .2 ft, and with additional pneumatic foramina (holes to allow air from air sacks to enter the bones) in the vertebrae and humerus. The describers concluded that the species was a member of the

Pterodactyloidea Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...

and within this group, using the phylogeny of

David Unwin David (; , "beloved one") (traditional spelling), , ''Dāwūd''; grc-koi, Δαυΐδ, Dauíd; la, Davidus, David; gez , ዳዊት, ''Dawit''; xcl, Դաւիթ, ''Dawitʿ''; cu, Давíдъ, ''Davidŭ''; possibly meaning "beloved one". w ...

, a member of a clade that is now known as Lophocratia.Harris, J.D., and Carpenter, K. (1996). A large pterodactyloid from the Morrison Formation (Late Jurassic) of Garden Park, Colorado. ''Neues Jahrbuch für Geologie und Paläontologie - Monatshefte'' 1996(8):473-484. It was regarded as a potentially valid genus in the most recent review of Morrison pterosaurs.King, L.R., Foster, J.R., and Scheetz, R.D. (2006). New pterosaur specimens from the Morrison Formation and a summary of the Late Jurassic pterosaur record of the Rocky Mountain region. In: Foster, J.R., and Lucas, S.G. (eds.). ''Paleontology and Geology of the Upper Morrison Formation''. New Mexico Museum of Natural History and Science Bulletin 36:149-161. ISSN 1524-4156.

Classification

The cladogram below shows a

phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

published by Longrich, Martill, and Andres in 2018. They recovered ''Kepodactylus'' as a basal member of the family

Ctenochasmatidae Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early C ...

References

{{Taxonbar, from=Q2218920 Ctenochasmatoids Late Jurassic pterosaurs of North America Morrison fauna Taxa named by Kenneth Carpenter Fossil taxa described in 1996

Classification

See also

References