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Gavialis Papuensis
''Ikanogavialis'' is an extinct genus of gavialid crocodilian. Fossils have been found in the Urumaco Formation in Urumaco, Venezuela and the Solimões Formation of Brazil. The strata from which remains are found are late Miocene in age, rather than Pliocene as was once thought. A possible member of this genus survived into the Late Holocene on Muyua or Woodlark Island in Papua New Guinea. Description ''Ikanogavialis'' had a dorsoventrally deep snout and a distinctive notch between the dentary and maxillary alveoli. The external nares projected anterodorsally from the rostrum. This can be seen as a plesiomorphic characteristic in crocodilians, but given that the earliest gavialoids possessed dorsally projecting external nares, this feature can be seen as having been a reversal from the gavialoid apomorphy back to the crocodilian plesiomorphy rather than having been directly obtained from an early crocodilian ancestor. Species The type species of ''Ikanogavialis'' is ''I. game ...
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Late Miocene
The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million years ago) to 5.333 Ma. The evolution of life The gibbons (family Hylobatidae) and orangutans (genus ''Pongo'') are the first groups to split from the line leading to the hominins, including humans, then gorillas (genus ''Gorilla''), and finally, chimpanzees and bonobos (genus ''Pan (genus), Pan''). The splitting date between hominin and chimpanzee lineages is placed by some between 4 to 8 million years ago, that is, during the Late Miocene. References External links GeoWhen Database - Late Miocene
Miocene, .03 Miocene geochronology, 03 Messinian, * Tortonian, * {{geochronology-stub ...
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Papua New Guinea
Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country in Oceania that comprises the eastern half of the island of New Guinea and its offshore islands in Melanesia (a region of the southwestern Pacific Ocean north of Australia). Its capital, located along its southeastern coast, is Port Moresby. The country is the world's third largest island country, with an area of . At the national level, after being ruled by three external powers since 1884, including nearly 60 years of Australian administration starting during World War I, Papua New Guinea established its sovereignty in 1975. It became an independent Commonwealth realm in 1975 with Elizabeth II as its queen. It also became a member of the Commonwealth of Nations in its own right. There are 839 known languages of Papua New Guinea, one of ...
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Caiman
A caiman (also cayman as a variant spelling) is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico, Central and South America from marshes and swamps to mangrove rivers and lakes. They have scaly skin and live a fairly nocturnal existence. They are relatively small-sized crocodilians with an average maximum weight of depending on species, with the exception of the black caiman (''Melanosuchus niger''), which can grow more than in length and weigh in excess of 1,000 kg (2,200 Ib). The black caiman is the largest caiman species in the world and is found in the slow-moving rivers and lakes that surround the Amazon basin. The smallest species is the Cuvier's dwarf caiman (''Paleosuchus palpebrosus''), which grows to long. There are six different species of caiman found throughout the watery jungle habitats of Central and Southern America. The average length for most ...
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Gavialoids
Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are alive today, with ''Hanyusuchus'' having become extinct in the last few centuries. Extinct South American gavialoids likely dispersed in the mid Tertiary from Africa and Asia. Fossil remains of the Puerto Rican gavialoid '' Aktiogavialis puertorisensis'' were discovered in a cave located in San Sebastián, Puerto Rico and dated to the Oligocene. This individual is thought to have crossed the Atlantic coming from Africa, indicating that this species was able to withstand saltwater. Classification Gavialoidea is cladistically defined as ''Gavialis gangeticus'' (the gharial) and all crocodylians closer to it than to ''Alligator mississippiensis'' (the American alligator) or ''Crocodylus niloticus'' (the Nile crocodile). This is a stem-bas ...
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Piscogavialis
''Piscogavialis'' is an extinct monospecific genus of gryposuchine gavialid crocodylian. The only species yet known is ''P. jugaliperforatus''. Fossils of ''Piscogavialis'' have been found from the Mio-Pliocene Pisco Formation of the Sacaco Basin in southern Peru in 1998,''Piscogavialis''
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where it coexisted with the much smaller gavialid '' Sacacosuchus''. ''Piscogavialis'' is known only from a single specimen, but it represents some of the best preserved gavialid material known from



Siquisiquesuchus
''Siquisiquesuchus'' (meaning " Siquisique crocodile" after the town in Lara, Venezuela, near where the first described specimens were found) is an extinct genus of gavialid crocodilian. It is known from cranial remains and a few postcranial bones found in Miocene-age rocks of the Castillo Formation in northwestern Venezuela. Description ''Siquisiquesuchus'' is based on the holotype MBLUZ–P–5050, a nearly complete skull and lower jaws. It was found near Lara in rocks of the Early Miocene-age Castillo Formation. Two other partial skulls, partial vertebrae, a thigh bone, partial upper arm, and partial shin bone were recovered from another locality. These bones were described by Christopher Brochu and Ascanio Rincón in 2004. The type species is ''S. venezuelensis''. Like other gavialoids, ''Siquisiquesuchus'' had a long, narrow rostrum on its skull, accounting for approximately 60% of the skull's length. The number of teeth in the premaxillary bones at the tip of the snou ...
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Gryposuchus
''Gryposuchus'' is an extinct genus of gavialid crocodilian. Fossils have been found from Argentina, Colombia, Venezuela, Brazil and the Peruvian Amazon. The genus existed during the Miocene epoch (Colhuehuapian to Huayquerian). One recently described species, ''G. croizati'', grew to an estimated length of . ''Gryposuchus'' is the type genus of the subfamily Gryposuchinae, although a 2018 study indicates that Gryposuchinae and ''Gryposuchus'' might be paraphyletic and rather an evolutionary grade towards the gharial. Species The type species of ''Gryposuchus'' is ''G. jessei'', named based on a well-preserved rostrum collected along the Pauini River of Brazil in 1912. The specimen was probably destroyed during World War II by the 1943 bombing of Hamburg. Another specimen named UFAC 1272, consisting of a premaxilla and maxilla, was discovered in the nearby Sena Madureia locality of the late Miocene Solimões Formation, in and referred to the species in 1997. ''G. jesse ...
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PeerJ
''PeerJ'' is an open access peer-reviewed scientific mega journal covering research in the biological and medical sciences. It is published by a company of the same name that was co-founded by CEO Jason Hoyt (formerly at Mendeley) and publisher Peter Binfield (formerly at '' PLOS One''), with initial financial backing of US$950,000 from O'Reilly Media's O'Reilly AlphaTech Ventures, and later funding from Sage Publishing. PeerJ officially launched in June 2012, started accepting submissions on December 3, 2012, and published its first articles on February 12, 2013. The company is a member of CrossRef, CLOCKSS, ORCID, and the Open Access Scholarly Publishers Association. The company's offices are in Corte Madera (California, USA), and London (Great Britain). Submitted research is judged solely on scientific and methodological soundness (as at '' PLoS ONE''), with a facility for peer reviews to be published alongside each paper. Business model ''PeerJ'' uses a business model th ...
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Pleistocene
The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology. The name is a combination of Ancient Greek grc, label=none, πλεῖστος, pleīstos, most and grc, label=none, καινός, kainós (latinized as ), 'new'. At the end of the preceding Pliocene, the previously isolated North and South American continents were joined by the Isthmus of Panama, causing Great American Interchang ...
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Urumaco Formation
The Urumaco Formation is a formation in Venezuela that includes deposits from the Late Miocene. It is the site of several "giant forms": the turtles, crocodiles, sloths and rodents of Urumaco are among the largest of their groups. Location The Urumaco formation is located in the Urumaco region in the Caribbean coastal Falcón state. The deposits date from 10 to 5.3 million years ago and the Urumaco formation was deposited in an area with large rivers, swamps, estuaries, lagoons and shallow coastal seas. These conditions in the Late Miocene contrast strongly with the current dry environment in the area today. Fauna Cartilaginous fish There are 21 known species of cartilaginous fishes from the Urumaco Formation, belonging to the orders Lamniformes, Carcharhiniformes, Myliobatiformes and Rajiformes. ''Carcharhinus caquetius'' is an endemic species of predator shark from Urumaco. A large number of well-preserved fossils of the sawfish ''Pristis rostra'' have been found in th ...
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Apomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis ...
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