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Piscogavialis
''Piscogavialis'' is an extinct monospecific genus of gryposuchine gavialid crocodylian. The only species yet known is ''P. jugaliperforatus''. Fossils of ''Piscogavialis'' have been found from the Mio-Pliocene Pisco Formation of the Sacaco Basin in southern Peru in 1998,''Piscogavialis''
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where it coexisted with the much smaller gavialid '' Sacacosuchus''. ''Piscogavialis'' is known only from a single specimen, but it represents some of the best preserved gavialid material known from

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Sacacosuchus
''Sacacosuchus'' is an extinct monospecific genus of marine gavialid that lived along the coast of the south-east Pacific from approximately 19 to 6.3 million years ago. Its fossils have been found in the Chilcatay and Pisco Formations of Peru, where it coexisted with the much larger ''Piscogavialis''. Based on its skull, ''Sacacosuchus'' was most likely a generalist feeder with an estimated total body length of . Its extinction is thought to have been caused by a combination of factors including falling sea levels and global cooling. History and naming ''Sacacosuchus'' is known from several adult to juvenile specimens mostly stemming from the Sacaco locality of the Pisco Formation. The holotype represents a nearly complete skull, while some of the referred specimens include partial skulls and dentary bones. Other material was found in the Ica-Pisco Valley and the older Chilcatay Formation. The generic name derives from the Sacaco area of the Pisco Basin and the Greek "suchu ...
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Gryposuchinae
Gryposuchinae is an extinct subfamily (biology), subfamily of Gavialidae, gavialid crocodylians. Gryposuchines lived mainly in the Miocene of South America. However, ''Ikanogavialis, "Ikanogavialis" papuensis'' may have survived more recently, into the Late Pleistocene/Holocene. Most were long-snouted coastal forms. The group was named in 2007 and includes genera such as ''Gryposuchus'' and ''Aktiogavialis'', although a 2018 study indicates that the group might be paraphyletic and rather an evolutionary grade towards the gharial. Description Gryposuchines have long, narrow snouts and protruding eye sockets. One distinguishing feature of the group is the lack of a large exposure of the prootic bone around the trigeminal foramen, a hole in the side of the braincase wall. Classification Gryposuchinae was named in 2007 as a subfamily of closely related gavialid crocodilians. It was cladistics, cladistically defined as a stem-based taxon including ''Gryposuchus jessei'' and all croco ...
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Pisco Formation
The Pisco Formation is a geologic formation located in Peru, on the southern coastal desert of Ica, Peru, Ica and Arequipa. The approximately thick formation was deposited in the Pisco Basin, spanning an age from the Miocene, Middle Miocene up to the Early Pleistocene, roughly from 15 to 2 Ma. The tuffaceous sandstones, diatomaceous siltstones, Conglomerate (geology), conglomerates and dolomite (rock), dolomites were deposited in a lagoonal to near-shore depositional environment, environment, in bays similar to other Pacific South American formations as the Caldera Basin, Bahía Inglesa and Coquimbo Formations of Chile. Several specialists consider the Pisco Formation one of the most important Lagerstätten,Brand et al., 2004Brand et al., 2011 based on the large amount of exceptionally preserved marine fossils, including sharks (most notably megalodon), penguins, whales, dolphins, birds, marine crocodiles and aquatic giant sloths. Famous fossils found in these layers include the ...
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Gavialidae
Gavialidae is a family (biology), family of large semiaquatic crocodilians with elongated, narrow snouts. Gavialidae consists of two extant taxon, living species, the gharial (''Gavialis gangeticus'') and the false gharial (''Tomistoma schlegelii''), both occurring in Asia. Many extinct members are known from a broader range, including the recently extinct ''Hanyusuchus''. Gavialids are generally regarded as lacking the jaw strength to capture the large mammalian prey favoured by crocodiles and alligators of similar size so their thin snout is best used to catch fish, however the false gharial has been found to have a Generalist and specialist species, generalist diet with mature adults preying upon larger vertebrates, such as Ungulate, ungulates. Taxonomy The family Gavialidae was proposed by Arthur Adams (zoologist), Arthur Adams in 1854 for reptiles with a very long and slender muzzle, webbed feet and nearly equal teeth. It is currently recognized as a crown group, meaning that ...
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Thalassocnus
''Thalassocnus'' is an extinct genus of semiaquatic ground sloths from the Miocene and Pliocene of the Pacific South American coast. It is monotypic within the subfamily Thalassocninae. The five species—''T. antiquus'', ''T. natans'', ''T. littoralis'', ''T. carolomartini'', and ''T. yuacensis''—represent a chronospecies, a population gradually adapting to marine life in one direct lineage. They are the only known aquatic sloths, but they may have also been adapted to a terrestrial lifestyle. They have been found in the Pisco Formation of Peru, the Tafna Formation of Argentina, and the Bahía Inglesa, Coquimbo, and Horcón formations of Chile. Thalassocninae has been placed in both the families Megatheriidae and Nothrotheriidae. ''Thalassocnus'' evolved several marine adaptations over the course of 4 million years, such as dense and heavy bones to counteract buoyancy, the internal nostrils migrating farther into the head to help with breathing while completely submerged, ...
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Gharial
The gharial (''Gavialis gangeticus''), also known as gavial or fish-eating crocodile, is a crocodilian in the family Gavialidae and among the longest of all living crocodilians. Mature females are long, and males . Adult males have a distinct boss at the end of the snout, which resembles an earthenware pot known as a ''ghara'', hence the name "gharial". The gharial is well adapted to catching fish because of its long, narrow snout and 110 sharp, interlocking teeth. The gharial probably evolved in the northern Indian subcontinent. Fossil gharial remains were excavated in Pliocene deposits in the Sivalik Hills and the Narmada River valley. It currently inhabits rivers in the plains of the northern part of the Indian subcontinent. It is the most thoroughly aquatic crocodilian, and leaves the water only for basking and building nests on moist sandbanks. Adults mate at the end of the cold season. Females congregate in spring to dig nests, in which they lay 20–95 eggs. They guard t ...
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False Gharial
The false gharial (''Tomistoma schlegelii''), also known by the names Malayan gharial, Sunda gharial and tomistoma is a freshwater crocodilian of the family Gavialidae native to Peninsular Malaysia, Borneo, Sumatra and Java. It is listed as Vulnerable on the IUCN Red List, as the global population is estimated at around 2,500 to 10,000 mature individuals. The specific name ''schlegelii'' honors Hermann Schlegel. Taxonomy The scientific name ''Crocodilus (Gavialis) schlegelii'' was proposed by Salomon Müller in 1838 who described a specimen collected in Borneo. In 1846, he proposed to use the name ''Tomistoma schlegelii'', if it needs to be placed in a distinct genus. The genus ''Tomistoma'' potentially also contains several extinct species like ''T. cairense'', ''T. lusitanicum'', ''T. taiwanicus'', and ''T. coppensi''. However, these species may need to be reclassified to different genera as are paraphyletic. The false gharial's snout broadens considerably towards the ...
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Late Miocene
The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million years ago) to 5.333 Ma. The evolution of life The gibbons (family Hylobatidae) and orangutans (genus ''Pongo'') are the first groups to split from the line leading to the hominins, including humans, then gorillas (genus ''Gorilla''), and finally, chimpanzees and bonobos (genus ''Pan (genus), Pan''). The splitting date between hominin and chimpanzee lineages is placed by some between 4 to 8 million years ago, that is, during the Late Miocene. References External links GeoWhen Database - Late Miocene
Miocene, .03 Miocene geochronology, 03 Messinian, * Tortonian, * {{geochronology-stub ...
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Extant Taxon
Neontology is a part of biology that, in contrast to paleontology, deals with living (or, more generally, '' recent'') organisms. It is the study of extant taxa (singular: extant taxon): taxa (such as species, genera and families) with members still alive, as opposed to (all) being extinct. For example: * The moose (''Alces alces'') is an extant species, and the dodo (''Raphus cucullatus'') is an extinct species. * In the group of molluscs known as the cephalopods, there were approximately 600 extant species and 7,500 extinct species. A taxon can be classified as extinct if it is broadly agreed or certified that no members of the group are still alive. Conversely, an extinct taxon can be reclassified as extant if there are new discoveries of living species ("Lazarus species"), or if previously-known extant species are reclassified as members of the taxon. Most biologists, zoologists, and botanists are in practice neontologists, and the term neontologist is used large ...
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Scientifically Described
A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have been described previously or are related. In order for species to be validly described, they need to follow guidelines established over time. Zoological naming requires adherence to the ICZN code, plants, the ICN, viruses ICTV, and so on. The species description often contains photographs or other illustrations of type material along with a note on where they are deposited. The publication in which the species is described gives the new species a formal scientific name. Some 1.9 million species have been identified and described, out of some 8.7 million that may actually exist. Millions more have become extinct throughout the existence of life on Earth. Naming process A name of a new species becomes valid (available in zool ...
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Marine Transgression
A marine transgression is a geologic event during which sea level rises relative to the land and the shoreline moves toward higher ground, which results in flooding. Transgressions can be caused by the land sinking or by the ocean basins filling with water or decreasing in capacity. Transgressions and regressions may be caused by tectonic events such as orogenies, severe climate change such as ice ages or isostatic adjustments following removal of ice or sediment load. During the Cretaceous, seafloor spreading created a relatively shallow Atlantic basin at the expense of deeper Pacific basin. That reduced the world's ocean basin capacity and caused a rise in sea level worldwide. As a result of the sea level rise, the oceans transgressed completely across the central portion of North America and created the Western Interior Seaway from the Gulf of Mexico to the Arctic Ocean. The opposite of transgression is regression in which the sea level falls relative to the land and expo ...
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Pisco Basin
Pisco Basin ( es, Cuenca de Pisco) is a sedimentary basin extending over in southwestern Peru.Solís Mundaca, 2018, p.1 The basin has a thick sedimentary fill, which is about half the thickness of more northern foreland basins in Peru. The oldest known sediments are the Eocene sandstones of the Caballas Formation, while the youngest deposits, the fossiliferous Pisco Formation, date to the Early Pleistocene. In relation to present-day, topography the fill of Pisco Basin makes the upper part of the Coastal Cordillera of southern Peru, the coastal plains, the Ica-Nazca Depression and the Andean foothills. The basin is renowned for hosting various highly fossiliferous stratigraphic units; the Pisco Formation has provided a wealth of marine mammals (including sloths), birds, fish and other groups, as have the Chilcatay, Otuma and Yumaque Formations. Stratigraphy Tectonic and sedimentary evolution The basin developed in a setting of extensional tectonics from Eocene to ...
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