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Siquisiquesuchus
''Siquisiquesuchus'' (meaning " Siquisique crocodile" after the town in Lara, Venezuela, near where the first described specimens were found) is an extinct genus of gavialid crocodilian. It is known from cranial remains and a few postcranial bones found in Miocene-age rocks of the Castillo Formation in northwestern Venezuela. Description ''Siquisiquesuchus'' is based on the holotype MBLUZ–P–5050, a nearly complete skull and lower jaws. It was found near Lara in rocks of the Early Miocene-age Castillo Formation. Two other partial skulls, partial vertebrae, a thigh bone, partial upper arm, and partial shin bone were recovered from another locality. These bones were described by Christopher Brochu and Ascanio Rincón in 2004. The type species is ''S. venezuelensis''. Like other gavialoids, ''Siquisiquesuchus'' had a long, narrow rostrum on its skull, accounting for approximately 60% of the skull's length. The number of teeth in the premaxillary bones at the tip of the snou ...
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Gryposuchinae
Gryposuchinae is an extinct subfamily (biology), subfamily of Gavialidae, gavialid crocodylians. Gryposuchines lived mainly in the Miocene of South America. However, ''Ikanogavialis, "Ikanogavialis" papuensis'' may have survived more recently, into the Late Pleistocene/Holocene. Most were long-snouted coastal forms. The group was named in 2007 and includes genera such as ''Gryposuchus'' and ''Aktiogavialis'', although a 2018 study indicates that the group might be paraphyletic and rather an evolutionary grade towards the gharial. Description Gryposuchines have long, narrow snouts and protruding eye sockets. One distinguishing feature of the group is the lack of a large exposure of the prootic bone around the trigeminal foramen, a hole in the side of the braincase wall. Classification Gryposuchinae was named in 2007 as a subfamily of closely related gavialid crocodilians. It was cladistics, cladistically defined as a stem-based taxon including ''Gryposuchus jessei'' and all croco ...
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Piscogavialis
''Piscogavialis'' is an extinct monospecific genus of gryposuchine gavialid crocodylian. The only species yet known is ''P. jugaliperforatus''. Fossils of ''Piscogavialis'' have been found from the Mio-Pliocene Pisco Formation of the Sacaco Basin in southern Peru in 1998,''Piscogavialis''
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where it coexisted with the much smaller gavialid '' Sacacosuchus''. ''Piscogavialis'' is known only from a single specimen, but it represents some of the best preserved gavialid material known from

Ikanogavialis
''Ikanogavialis'' is an extinct genus of gavialid crocodilian. Fossils have been found in the Urumaco Formation in Urumaco, Venezuela and the Solimões Formation of Brazil. The strata from which remains are found are late Miocene in age, rather than Pliocene as was once thought. A possible member of this genus survived into the Late Holocene on Muyua or Woodlark Island in Papua New Guinea. Description ''Ikanogavialis'' had a dorsoventrally deep snout and a distinctive notch between the dentary and maxillary alveoli. The external nares projected anterodorsally from the rostrum. This can be seen as a plesiomorphic characteristic in crocodilians, but given that the earliest gavialoids possessed dorsally projecting external nares, this feature can be seen as having been a reversal from the gavialoid apomorphy back to the crocodilian plesiomorphy rather than having been directly obtained from an early crocodilian ancestor. Species The type species of ''Ikanogavialis'' is ''I. ga ...
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Gryposuchus
''Gryposuchus'' is an extinct genus of gavialid crocodilian. Fossils have been found from Argentina, Colombia, Venezuela, Brazil and the Peruvian Amazon. The genus existed during the Miocene epoch (Colhuehuapian to Huayquerian). One recently described species, ''G. croizati'', grew to an estimated length of . ''Gryposuchus'' is the type genus of the subfamily Gryposuchinae, although a 2018 study indicates that Gryposuchinae and ''Gryposuchus'' might be paraphyletic and rather an evolutionary grade towards the gharial. Species The type species of ''Gryposuchus'' is ''G. jessei'', named based on a well-preserved rostrum collected along the Pauini River of Brazil in 1912. The specimen was probably destroyed during World War II by the 1943 bombing of Hamburg. Another specimen named UFAC 1272, consisting of a premaxilla and maxilla, was discovered in the nearby Sena Madureia locality of the late Miocene Solimões Formation, in and referred to the species in 1997. ''G. jesse ...
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Gavialidae
Gavialidae is a family (biology), family of large semiaquatic crocodilians with elongated, narrow snouts. Gavialidae consists of two extant taxon, living species, the gharial (''Gavialis gangeticus'') and the false gharial (''Tomistoma schlegelii''), both occurring in Asia. Many extinct members are known from a broader range, including the recently extinct ''Hanyusuchus''. Gavialids are generally regarded as lacking the jaw strength to capture the large mammalian prey favoured by crocodiles and alligators of similar size so their thin snout is best used to catch fish, however the false gharial has been found to have a Generalist and specialist species, generalist diet with mature adults preying upon larger vertebrates, such as Ungulate, ungulates. Taxonomy The family Gavialidae was proposed by Arthur Adams (zoologist), Arthur Adams in 1854 for reptiles with a very long and slender muzzle, webbed feet and nearly equal teeth. It is currently recognized as a crown group, meaning that ...
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Gavialid
Gavialidae is a family of large semiaquatic crocodilians with elongated, narrow snouts. Gavialidae consists of two living species, the gharial (''Gavialis gangeticus'') and the false gharial (''Tomistoma schlegelii''), both occurring in Asia. Many extinct members are known from a broader range, including the recently extinct ''Hanyusuchus''. Gavialids are generally regarded as lacking the jaw strength to capture the large mammalian prey favoured by crocodiles and alligators of similar size so their thin snout is best used to catch fish, however the false gharial has been found to have a generalist diet with mature adults preying upon larger vertebrates, such as ungulates. Taxonomy The family Gavialidae was proposed by Arthur Adams in 1854 for reptiles with a very long and slender muzzle, webbed feet and nearly equal teeth. It is currently recognized as a crown group, meaning that it only includes the last common ancestor of all extant (living) gavialids (the gharial and false gha ...
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Castillo Formation, Venezuela
The Castillo Formation is an Early Miocene (Burdigalian, Colhuehuapian to Santacrucian in the SALMA classification) geologic formation in the Falcón Basin of Venezuela. The formation unconformably overlies the Matatere, Misoa, El Paují and Jarillal Formations.Urbani & Mendi, 2010, p.17 The Castillo Formation is overlain by Quaternary alluvium and in places by the Capadare Formation.Urbani & Mendi, 2010, p.19 The formation, deposited in a calm near-shore lagoonal brackish environment, with possibly fluvial influence, has provided a rich assemblage of fossil crocodylians, turtles, giant sloths and various types of fish. Description The Castillo Formation crops out cover a wide semicircular area that extends through the northwestern Venezuelan states of Falcón and Lara. During Oligocene to Miocene times, the formation formed the northwest to southeast edge of the Falcón Basin.Solórzano et al., 2018a, p.3 The formation, with a minimum thickness of ,Rincón et al., 2014, ...
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Tibia
The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects the knee with the ankle. The tibia is found on the medial side of the leg next to the fibula and closer to the median plane. The tibia is connected to the fibula by the interosseous membrane of leg, forming a type of fibrous joint called a syndesmosis with very little movement. The tibia is named for the flute ''tibia''. It is the second largest bone in the human body, after the femur. The leg bones are the strongest long bones as they support the rest of the body. Structure In human anatomy, the tibia is the second largest bone next to the femur. As in other vertebrates the tibia is one of two bones in the lower leg, the other being the fibula, and is a component of the knee and ankle joints. The ossification or formation of the bone ...
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Gavialis
''Gavialis'' is a genus of crocodylians that includes the living gharial ''Gavialis gangeticus'' and one known extinct species, '' Gavialis bengawanicus.'' ''G. gangeticus'' comes from the Indian Subcontinent, while ''G. bengawanicus'' is known from Java. ''Gavialis'' likely first appeared in the Indian Subcontinent in the Pliocene and dispersed into the Malay Archipelago through a path called the Siva–Malayan route in the Quaternary. Remains attributed to ''Gavialis'' have also been found on Sulawesi and Woodlark Island east of the Wallace Line, suggesting a prehistoric lineage of ''Gavialis'' was able to traverse marine environments and reach places possibly as far as western Oceania. The genus ''Gavialis'' was reevaluated in 2018 based on specimens in the Natural History Museum, London that were collected in the Sivalik Hills. The author concluded that ''G. gangeticus'' and '' G. bengawanicus'' are the only two species in the genus ''Gavialis'', with '' G. hysudricus'' a ...
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Suture (anatomical)
In anatomy, a suture is a fairly rigid joint between two or more hard elements of an organism, with or without significant overlap of the elements. Sutures are found in the skeletons or exoskeletons of a wide range of animals, in both invertebrates and vertebrates. Sutures are found in animals with hard parts from the Cambrian period to the present day. Sutures were and are formed by several different methods, and they exist between hard parts that are made from several different materials. Vertebrate skeletons The skeletons of vertebrate animals (fish, amphibians, reptiles, birds, and mammals) are made of bone, in which the main rigid ingredient is calcium phosphate. Cranial sutures The skulls of most vertebrates consist of sets of bony plates held together by cranial sutures. These sutures are held together mainly by Sharpey's fibers which grow from each bone into the adjoining one. Sutures in the ankles of land vertebrates In the type of crurotarsal ankle which is found i ...
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Dentary
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movable bone of the skull (discounting the ossicles of the middle ear). It is connected to the temporal bones by the temporomandibular joints. The bone is formed prenatal development, in the fetus from a fusion of the left and right mandibular prominences, and the point where these sides join, the mandibular symphysis, is still visible as a faint ridge in the midline. Like other symphyses in the body, this is a midline articulation where the bones are joined by fibrocartilage, but this articulation fuses together in early childhood.Illustrated Anatomy of the Head and Neck, Fehrenbach and Herring, Elsevier, 2012, p. 59 The word "mandible" derives from the Latin word ''mandibula'', "jawbone" (literally "one used for chewing"), from ''wikt:mandere ...
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Maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the mandibular symphysis. The mandible is the movable part of the jaw. Structure In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the alveolar process ** the palatine process * three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen Articulations Each maxilla articulates with nine bones: * two of the cranium: the frontal and ethmoid * seven of the face: the nasal, zygomatic, lacrimal, inferior n ...
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