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Gallardosaurus
''Gallardosaurus'' is a genus of pliosaurid plesiosaur from the Caribbean seaway. It contains the single species ''Gallardosaurus iturraldei''. ''Gallardosaurus'' was found in middle-late Oxfordian-age (Late Jurassic) rocks of the Jagua Formation of western Cuba.''Gallardosaurus''
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''Gallardosaurus'' is believed to be evolutionarily connected to '''', a pliosaurid commonly found in the Oxfordian-aged sediment.


History and etymology


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Gallardosaurus Skeletal
''Gallardosaurus'' is a genus of pliosaurid plesiosaur from the Caribbean seaway. It contains the single species ''Gallardosaurus iturraldei''. ''Gallardosaurus'' was found in middle-late Oxfordian-age (Late Jurassic) rocks of the Jagua Formation of western Cuba.''Gallardosaurus''
at .org
''Gallardosaurus'' is believed to be evolutionarily connected to '''', a pliosaurid commonly found in the Oxfordian-aged sediment.


History and etymology


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Peloneustes
''Peloneustes'' (meaning "mud swimmer") is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of ''Plesiosaurus'' by palaeontologist Harry Govier Seeley in 1896, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to ''Peloneustes'', ''P. philarchus'' is currently the only one still considered valid, with the others moved to different genera, considered '' nomina dubia'', or synonymised with ''P. philarchus''. Some of the material formerly assigned to ''P. evansi'' have since been reassigned to ''"Pliosaurus" andrewsi''. ''Peloneustes'' is known from many specimens, including some very complete material. With a total length of , ''Peloneustes'' is not a large pliosaurid. It had a large, triangular skull, which occupied about a fifth of its body length. The ...
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Pliosaurus
''Pliosaurus'' (meaning 'more lizard') is an extinct genus of thalassophonean pliosaurid known from the Kimmeridgian and Tithonian stages (Late Jurassic) of Europe and South America. Their diet would have included fish, cephalopods, and marine reptiles. This genus has contained many species in the past but recent reviews found only six (''P. brachydeirus'', ''P. carpenteri'', ''P. funkei'', ''P. kevani'', ''P. rossicus'' and ''P. westburyensis'') to be valid, while the validity of two additional species awaits a petition to the International Code of Zoological Nomenclature. Currently, ''P. brachyspondylus'' and ''P. macromerus'' are considered dubious, while ''P. portentificus'' is considered undiagnostic. Most species of ''Pliosaurus'' reached in length and in body mass, while ''P. rossicus'' and ''P. funkei'' may have reached or even exceeded in length and in body mass, being the largest plesiosaurs of all time. Species of this genus are differentiated from other pliosaurids ...
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Jagua Formation
The Jagua Formation is a Late Jurassic (middle to late Oxfordian) geologic formation in the Sierra de los Órganos and Sierra del Rosario mountain ranges in Pinar del Río Province, western Cuba. Plesiosaur, pliosaur, pterosaur,Barrett et al., 2008 metriorhynchid, turtle and dinosaur remains are among the fossils that have been recovered from its strata. Description The formation comprises marine shales and limestones. The thick Jagua Vieja Member consists of black shales and horizontally laminated marly micritic to biomicritic limestones. The latter contains calcareous concretions which the fossils are found in. The formation overlies the San Cayetano Formation and is overlain by the Guasasa Formation. The bedding direction is steeply dipping towards the northwest. Fossil content Vertebrate paleofauna Other fossils *'' Favreina'' *'' Globochaete'' * '' Gryphaea mexicana'' - Pan de Azúcar Mb. * '' Lepidotes gloriae'' * '' Liostrea mairei'' * '' L. sandalina'' ...
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Plesiosauria
The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period, possibly in the Rhaetian stage, about 203 million years ago. They became especially common during the Jurassic Period, thriving until their disappearance due to the Cretaceous–Paleogene extinction event at the end of the Cretaceous Period, about 66 million years ago. They had a worldwide oceanic distribution, and some species at least partly inhabited freshwater environments. Plesiosaurs were among the first fossil reptiles discovered. In the beginning of the nineteenth century, scientists realised how distinctive their build was and they were named as a separate order in 1835. The first plesiosaurian genus, the eponymous ''Plesiosaurus'', was named in 1821. Since then, more than a hundred vali ...
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Pliosauridae
Pliosauridae is a family of plesiosaurian marine reptiles from the Latest Triassic to the early Late Cretaceous ( Rhaetian to Turonian stages) of Australia, Europe, North America and South America. The family is more inclusive than the archetypal short-necked large headed species that are placed in the subclade Thalassophonea, with basal forms resembling other plesiosaurs with long necks. They became extinct during the early Late Cretaceous and were subsequently replaced by the mosasaurs. It was formally named by Harry G. Seeley in 1874. Relationships Pliosauridae is a stem-based taxon defined in 2010 (and in earlier studies in a similar manner) as "all taxa more closely related to ''Pliosaurus brachydeirus'' than to '' Leptocleidus superstes'', '' Polycotylus latipinnis'' or ''Meyerasaurus victor''". The family Brachauchenidae has been proposed to include pliosauroids which have very short necks and may include ''Brachauchenius'' and ''Kronosaurus''. However, modern cladisti ...
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Liopleurodon
''Liopleurodon'' (; meaning 'smooth-sided teeth') is an extinct genus of large, carnivorous marine reptile belonging to the Thalassophonea, a clade of short-necked pliosaurid plesiosaurs. ''Liopleurodon'' lived from the Callovian Stage of the Middle Jurassic to the Kimmeridgian stage of the Late Jurassic Period (c. 166 to 155 mya). It was the apex predator of the Middle to Late Jurassic seas that covered Europe. The largest species, ''L. ferox'', is estimated to have grown up to in length, but could have been larger. The name "Liopleurodon" (meaning "smooth-sided tooth") derives from Ancient Greek words: ', "smooth"; ', "side" or "rib"; and ', "tooth". Discovery and species Even before ''Liopleurodon'' was named, material likely belonging to it was described. In 1841, Hermann von Meyer named the species ''Thaumatosaurus oolithicus'' based on a fragmentary specimen consisting of partial teeth, skull elements, vertebrae, and ribs from deposits in Württemberg, Germany poss ...
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Pliosaur
Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled pleisiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used. The distinguishing characteristics are a short neck and an elongated head, with larger hind flippers compared to the fore flippers, the opposite of the plesiosaurs. They were carnivorous and their long and powerful jaws carried many sharp, conical teeth. Pliosaurs range from 4 to 15 metres and more in length. Their prey may have included fish, sharks, i ...
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Plesiosaur
The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period, possibly in the Rhaetian stage, about 203 million years ago. They became especially common during the Jurassic Period, thriving until their disappearance due to the Cretaceous–Paleogene extinction event at the end of the Cretaceous Period, about 66 million years ago. They had a worldwide oceanic distribution, and some species at least partly inhabited freshwater environments. Plesiosaurs were among the first fossil reptiles discovered. In the beginning of the nineteenth century, scientists realised how distinctive their build was and they were named as a separate order in 1835. The first plesiosaurian genus, the eponymous ''Plesiosaurus'', was named in 1821. Since then, more than a hundred valid ...
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Oxfordian Stage
The Oxfordian is, in the ICS' geologic timescale, the earliest age of the Late Jurassic Epoch, or the lowest stage of the Upper Jurassic Series. It spans the time between 163.5 ± 1.0 Ma and 157.3 ± 1.0 Ma (million years ago). The Oxfordian is preceded by the Callovian and is followed by the Kimmeridgian. Stratigraphic definitions The Oxfordian Stage was called "Clunch Clay and Shale" by William Smith (1815–1816); in 1818 W. Buckland described them under the unwieldy title "Oxford, Forest or Fen Clay". The term Oxfordian was introduced by Alcide d'Orbigny in 1844. The name is derived from the English city of Oxford, where the beds are well developed, but they crop out almost continuously from Dorset to the coast of Yorkshire, generally forming low, broad valleys. They are well exposed at Weymouth, Oxford, Bedford, Peterborough, and in the cliffs at Scarborough, Red Cliff and Gristhorpe Bay. Rocks of this age are found also in Uig and Skye. The base of the Oxfordian Stage i ...
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Pinar Del Río Province
Pinar del Río is one of the provinces of Cuba. It is at the western end of the island of Cuba. Geography The Pinar del Río province is Cuba's westernmost province and contains one of Cuba's three main mountain ranges, the Cordillera de Guaniguanico, divided into the easterly Sierra del Rosario and the westerly Sierra de los Órganos. These form a landscape characterised by steep sided limestone hills (called mogotes) and flat, fertile valleys. One such topographic feature, the Viñales Valley, is a UNESCO World Heritage Site. The northern coast opens to the great Gulf of Mexico, and is lined by the Colorados Archipelago, a string of cays and isles developed on a reef barrier. The westernmost point of Cuba, Cabo San Antonio, is located on the Guanahacabibes Peninsula, which is a National Park and a Biosphere Reserve. History The city was founded by the Spanish as ''Nueva Filipinas'' (New Philippines), and the city was renamed Pinar del Río in 1774. The province was founded i ...
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Concretion
A concretion is a hard, compact mass of matter formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes also occur. The word 'concretion' is derived from the Latin "(act of) compacting, condensing, congealing, uniting", itself from ''con'' meaning 'together' and ''crescere'' meaning "to grow". Concretions form within layers of sedimentary strata that have already been deposited. They usually form early in the burial history of the sediment, before the rest of the sediment is hardened into rock. This concretionary cement often makes the concretion harder and more resistant to weathering than the host stratum. There is an important distinction to draw between concretions and nodules. Concretions are formed from mineral precipitation around some kind of nucleus while a nodule is a replacement body. Descriptions dating from the 18th ce ...
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