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Peloneustes
''Peloneustes'' (meaning "mud swimmer") is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of ''Plesiosaurus'' by palaeontologist Harry Govier Seeley in 1896, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to ''Peloneustes'', ''P. philarchus'' is currently the only one still considered valid, with the others moved to different genera, considered '' nomina dubia'', or synonymised with ''P. philarchus''. Some of the material formerly assigned to ''P. evansi'' have since been reassigned to ''"Pliosaurus" andrewsi''. ''Peloneustes'' is known from many specimens, including some very complete material. With a total length of , ''Peloneustes'' is not a large pliosaurid. It had a large, triangular skull, which occupied about a fifth of its body length. The ...
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Peloneustes Holotype Rostrum
''Peloneustes'' (meaning "mud swimmer") is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of ''Plesiosaurus'' by palaeontologist Harry Govier Seeley in 1896, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to ''Peloneustes'', ''P. philarchus'' is currently the only one still considered valid, with the others moved to different genera, considered ''nomina dubia'', or synonymised with ''P. philarchus''. Some of the material formerly assigned to ''P. evansi'' have since been reassigned to ''"Pliosaurus" andrewsi''. ''Peloneustes'' is known from many specimens, including some very complete material. With a total length of , ''Peloneustes'' is not a large pliosaurid. It had a large, triangular skull, which occupied about a fifth of its body length. The ...
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Plesiosaur
The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period, possibly in the Rhaetian stage, about 203 million years ago. They became especially common during the Jurassic Period, thriving until their disappearance due to the Cretaceous–Paleogene extinction event at the end of the Cretaceous Period, about 66 million years ago. They had a worldwide oceanic distribution, and some species at least partly inhabited freshwater environments. Plesiosaurs were among the first fossil reptiles discovered. In the beginning of the nineteenth century, scientists realised how distinctive their build was and they were named as a separate order in 1835. The first plesiosaurian genus, the eponymous ''Plesiosaurus'', was named in 1821. Since then, more than a hundred valid ...
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Oxford Clay Formation
The Oxford Clay (or Oxford Clay Formation) is a Jurassic marine sedimentary rock formation underlying much of southeast England, from as far west as Dorset and as far north as Yorkshire. The Oxford Clay Formation dates to the Jurassic, specifically, the Callovian and Oxfordian ages, and comprises two main facies. The lower facies comprises the Peterborough Member, a fossiliferous organic-rich mudstone. This facies and its rocks are commonly known as lower Oxford Clay. The upper facies comprises the middle Oxford Clay, the Stewartby Member, and the upper Oxford Clay, the Weymouth Member. The upper facies is a fossil poor assemblage of calcareous mudstones. Oxford Clay appears at the surface around Oxford, Peterborough and Weymouth and is exposed in many quarries around these areas. The top of the Lower Oxford Clay shows a lithological change, where fissile shale changes to grey mudstone. The Middle and Upper Oxford Clays differ slightly, as they are separated by an argillaceou ...
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Thalassophonea
Thalassophonea is an extinct clade of pliosaurids from the Middle Jurassic to the early Late Cretaceous (Callovian to Turonian) of Australia, Europe, North America and South America. ''Thalassophonea'' was erected by Roger Benson and Patrick Druckenmiller in 2013. The name is derived from Greek ''thalassa'' (θάλασσα), "sea", and ''phoneus'' (φονεύς), "murderer". It is a stem-based taxon defined as "all taxa more closely related to ''Pliosaurus brachydeirus'' than to ''Marmornectes candrewi''". It includes the short necked and large headed taxa that typify the family. Classification The following cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... follows an analysis by Benson & Druckenmiller (2014). References Middle Jurassic first appearances C ...
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Pliosaurus
''Pliosaurus'' (meaning 'more lizard') is an extinct genus of thalassophonean pliosaurid known from the Kimmeridgian and Tithonian stages (Late Jurassic) of Europe and South America. Their diet would have included fish, cephalopods, and marine reptiles. This genus has contained many species in the past but recent reviews found only six (''P. brachydeirus'', ''P. carpenteri'', ''P. funkei'', ''P. kevani'', ''P. rossicus'' and ''P. westburyensis'') to be valid, while the validity of two additional species awaits a petition to the International Code of Zoological Nomenclature. Currently, ''P. brachyspondylus'' and ''P. macromerus'' are considered dubious, while ''P. portentificus'' is considered undiagnostic. Most species of ''Pliosaurus'' reached in length and in body mass, while ''P. rossicus'' and ''P. funkei'' may have reached or even exceeded in length and in body mass, being the largest plesiosaurs of all time. Species of this genus are differentiated from other pliosaurids ...
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Pliosaurid
Pliosauridae is a family of plesiosaurian marine reptiles from the Latest Triassic to the early Late Cretaceous ( Rhaetian to Turonian stages) of Australia, Europe, North America and South America. The family is more inclusive than the archetypal short-necked large headed species that are placed in the subclade Thalassophonea, with basal forms resembling other plesiosaurs with long necks. They became extinct during the early Late Cretaceous and were subsequently replaced by the mosasaurs. It was formally named by Harry G. Seeley in 1874. Relationships Pliosauridae is a stem-based taxon defined in 2010 (and in earlier studies in a similar manner) as "all taxa more closely related to ''Pliosaurus brachydeirus'' than to '' Leptocleidus superstes'', '' Polycotylus latipinnis'' or ''Meyerasaurus victor''". The family Brachauchenidae has been proposed to include pliosauroids which have very short necks and may include ''Brachauchenius'' and ''Kronosaurus''. However, modern cladisti ...
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Plesiosaurus
''Plesiosaurus'' (Greek: ' ('), near to + ' ('), lizard) is a genus of extinct, large marine sauropterygian reptile that lived during the Early Jurassic. It is known by nearly complete skeletons from the Lias Group, Lias of England. It is distinguishable by its small head, long and slender neck, broad turtle-like body, a short tail, and two pairs of large, elongated paddles. It lends its name to the order Plesiosauria, of which it is an early, but fairly typical member. It contains only one species, the type species, type, ''Plesiosaurus dolichodeirus''. Other species once assigned to this genus, including ''P. brachypterygius'', ''P. guilielmiimperatoris'', and ''P. tournemirensis'' have been reassigned to new genera, such as ''Hydrorion'', ''Seeleyosaurus'' and ''Occitanosaurus''. Discovery The first complete skeleton of ''Plesiosaurus'' was discovered by early paleontologist and fossil hunter Mary Anning in Sinemurian (Early Jurassic)-age rocks of the lower Lias Group in Dec ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Subaqueous Flight
Aquatic locomotion or swimming is biologically propelled motion through a liquid medium. The simplest propulsive systems are composed of cilia and flagella. Swimming has evolved a number of times in a range of organisms including arthropods, fish, molluscs, amphibians, reptiles, birds, and mammals. Evolution of swimming Swimming evolved a number of times in unrelated lineages. Supposed jellyfish fossils occur in the Ediacaran, but the first free-swimming animals appear in the Early to Middle Cambrian. These are mostly related to the arthropods, and include the Anomalocaridids, which swam by means of lateral lobes in a fashion reminiscent of today's cuttlefish. Cephalopods joined the ranks of the nekton in the late Cambrian, and chordates were probably swimming from the Early Cambrian. Many terrestrial animals retain some capacity to swim, however some have returned to the water and developed the capacities for aquatic locomotion. Most apes (including humans), however, lost th ...
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Fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts. The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period. Although they lacked a true spine, they possessed notochords which allowed them to be more agile than their invertebrate counterparts. Fish would continue to evolve through the Paleozoic era, diversifying into a wide variety of forms. Many fish of the Paleozoic developed external armor that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many (such as sharks) became formidable marine predators rather than just the prey of arthropods. Mos ...
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Epicontinental Sea
An inland sea (also known as an epeiric sea or an epicontinental sea) is a continental body of water which is very large and is either completely surrounded by dry land or connected to an ocean by a river, strait, or "arm of the sea". An inland sea will generally have higher salinity than a freshwater lake, but usually lower salinity than the open ocean. Definition What constitutes an "inland sea" is complex and somewhat necessarily vague. The United States Hydrographic Office defined it as "a body of water nearly or completely surrounded by land, especially if very large or composed of salt water". Geologic engineers Heinrich Ries and Thomas L. Watson say an inland sea is merely a very large lake. Rydén, Migula, and Andersson and Deborah Sandler of the Environmental Law Institute add that an inland sea is "more or less" cut off from the ocean. It may be semi-enclosed, or connected to the ocean by a strait or "arm of the sea". An inland sea is distinguishable from a bay in t ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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