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Pliosauridae is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
plesiosauria The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...
n marine reptiles from the Latest Triassic to the early
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
( Rhaetian to
Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded b ...
stages) of
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
. The family is more inclusive than the archetypal short-necked large headed species that are placed in the subclade
Thalassophonea Thalassophonea is an extinct clade of pliosaurids from the Middle Jurassic to the early Late Cretaceous ( Callovian to Turonian) of Australia, Europe, North America and South America. ''Thalassophonea'' was erected by Roger Benson and Patrick ...
, with basal forms resembling other plesiosaurs with long necks. They became extinct during the early
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
and were subsequently replaced by the
mosasaurs Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on th ...
. It was formally named by Harry G. Seeley in 1874.


Relationships

Pliosauridae is a
stem-based taxon Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which ...
defined in 2010 (and in earlier studies in a similar manner) as "all taxa more closely related to ''
Pliosaurus brachydeirus ''Pliosaurus'' (meaning 'more lizard') is an extinct genus of thalassophonean pliosaurid known from the Kimmeridgian and Tithonian stages (Late Jurassic) of Europe and South America. Their diet would have included fish, cephalopods, and marine re ...
'' than to '' Leptocleidus superstes'', '' Polycotylus latipinnis'' or ''
Meyerasaurus victor ''Meyerasaurus'' is an extinct genus of rhomaleosaurid known from Holzmaden, Baden-Württemberg of southwestern Germany. Description ''Meyerasaurus'' is known from the holotype SMNS 12478, articulated and complete skeleton which preserv ...
''". The family Brachauchenidae has been proposed to include pliosauroids which have very short necks and may include ''
Brachauchenius ''Brachauchenius'' (meaning 'short neck') is an extinct genus of pliosaurid that lived in North America (United States) and Morocco during the Late Cretaceous. History The type species, ''Brachauchenius lucasi'', lived in the Western Inland S ...
'' and ''
Kronosaurus ''Kronosaurus'' ( ; meaning "lizard of Kronos") is a potentially dubious genus of extinct short-necked pliosaur. With an estimated length of , it was among the largest pliosaurs, and is named after the leader of the Greek Titans, Kronos. It ...
''. However, modern
cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
analyses found that this group is actually a subfamily of pliosaurids, and possibly even the "crown group" of Pliosauridae. The following
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
follows an analysis by Benson & Druckenmiller (2014).