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Francevillian Biota
The Francevillian biota (also known as Gabon macrofossils or Gabonionta) is a group of 2.1-billion-year-old Palaeoproterozoic, macroscopic organisms known from fossils found in Gabon in the Palaeoproterozoic Francevillian B Formation, a black shale province. The fossils are postulated to be evidence of the earliest form of multicellular life. They were discovered by an international team led by the Moroccan-French geologist Abderrazak El Albani, of the University of Poitiers, France. While they have yet to be assigned to a formal taxonomic position, they have been informally and collectively referred to as the "Gabonionta" by the Natural History Museum Vienna in 2014. Morphology The fossil organisms are up to in size. Their bodies were flattened disks with a characteristic morphology, including circular and elongated individuals. A spherical to ellipsoidal central body is bounded by radial structures. The fossils show three-dimensionality and coordinated growth. Cell-cell comm ...
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Palaeoproterozoic
The Paleoproterozoic Era (;, also spelled Palaeoproterozoic), spanning the time period from (2.5–1.6  Ga), is the first of the three sub-divisions (eras) of the Proterozoic Eon. The Paleoproterozoic is also the longest era of the Earth's geological history. It was during this era that the continents first stabilized. Paleontological evidence suggests that the Earth's rotational rate ~1.8 billion years ago equated to 20-hour days, implying a total of ~450 days per year. Atmosphere Before the enormous increase in atmospheric oxygen, almost all existing lifeforms were anaerobic organisms whose metabolism was based on a form of cellular respiration that did not require oxygen. Free oxygen in large amounts is toxic to most anaerobic organisms. Consequently, most died when the atmospheric free oxygen levels soared in an extinction event called the Great Oxidation Event, which brought atmospheric oxygen levels to up to 10% of their current level. The only creatures that survi ...
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Francevillian Biota
The Francevillian biota (also known as Gabon macrofossils or Gabonionta) is a group of 2.1-billion-year-old Palaeoproterozoic, macroscopic organisms known from fossils found in Gabon in the Palaeoproterozoic Francevillian B Formation, a black shale province. The fossils are postulated to be evidence of the earliest form of multicellular life. They were discovered by an international team led by the Moroccan-French geologist Abderrazak El Albani, of the University of Poitiers, France. While they have yet to be assigned to a formal taxonomic position, they have been informally and collectively referred to as the "Gabonionta" by the Natural History Museum Vienna in 2014. Morphology The fossil organisms are up to in size. Their bodies were flattened disks with a characteristic morphology, including circular and elongated individuals. A spherical to ellipsoidal central body is bounded by radial structures. The fossils show three-dimensionality and coordinated growth. Cell-cell comm ...
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Cell-cell Communication
In biology, cell signaling (cell signalling in British English) or cell communication is the ability of a cell to receive, process, and transmit signals with its environment and with itself. Cell signaling is a fundamental property of all cellular life in prokaryotes and eukaryotes. Signals that originate from outside a cell (or extracellular signals) can be physical agents like mechanical pressure, voltage, temperature, light, or chemical signals (e.g., small molecules, peptides, or gas). Cell signaling can occur over short or long distances, and as a result can be classified as autocrine, juxtacrine, intracrine, paracrine, or endocrine. Signaling molecules can be synthesized from various biosynthetic pathways and released through passive or active transports, or even from cell damage. Receptors play a key role in cell signaling as they are able to detect chemical signals or physical stimuli. Receptors are generally proteins located on the cell surface or within the interior ...
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Microbial Mat
A microbial mat is a multi-layered sheet of microorganisms, mainly bacteria and archaea, or bacteria alone. Microbial mats grow at interfaces between different types of material, mostly on submerged or moist surfaces, but a few survive in deserts. A few are found as endosymbionts of animals. Although only a few centimetres thick at most, microbial mats create a wide range of internal chemical environments, and hence generally consist of layers of microorganisms that can feed on or at least tolerate the dominant chemicals at their level and which are usually of closely related species. In moist conditions mats are usually held together by slimy substances secreted by the microorganisms. In many cases some of the bacteria form tangled webs of filaments which make the mat tougher. The best known physical forms are flat mats and stubby pillars called stromatolites, but there are also spherical forms. Microbial mats are the earliest form of life on Earth for which there is good fossi ...
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Eukaryote
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard archaea. This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as flagellated phagotrophs. Their name comes from the Greek εὖ (''eu'', "well" or "good") and κάρυον (''karyon'', "nut" or "kernel"). Euka ...
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Aerobic Respiration
Cellular respiration is the process by which biological fuels are oxidised in the presence of an inorganic electron acceptor such as oxygen to produce large amounts of energy, to drive the bulk production of ATP. Cellular respiration may be described as a set of metabolic reactions and processes that take place in the cells of organisms to convert chemical energy from nutrients into adenosine triphosphate (ATP), and then release waste products. The reactions involved in respiration are catabolic reactions, which break large molecules into smaller ones, releasing energy. Respiration is one of the key ways a cell releases chemical energy to fuel cellular activity. The overall reaction occurs in a series of biochemical steps, some of which are redox reactions. Although cellular respiration is technically a combustion reaction, it is an unusual one because of the slow, controlled release of energy from the series of reactions. Nutrients that are commonly used by animal and pla ...
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Progradation
In sedimentary geology and geomorphology, the term progradation refers to the growth of a river delta farther out into the sea over time. This occurs when the volume of incoming sediment is greater than the volume of the delta that is lost through subsidence, sea-level rise, or erosion. Progradation can be caused by: * Periods of sea-level fall which result in marine regression. This can occur during major continental glaciations within ice ages, be caused by changes in the rates of seafloor spreading that affects the volume of the ocean basins, or tectonic effects on the regional mantle density structure that can change the geoid elevation. * Extremely high sediment input, such as by the Huang He (Yellow River) in China, which drains the Loess plateau, or from high sediment loads in proglacial river A proglacial river is a river that flows from the margin of a glacier. These rivers are strongly affected by the highly-seasonal water supply from the glacier and by the large supply ...
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Geochemistry
Geochemistry is the science that uses the tools and principles of chemistry to explain the mechanisms behind major geological systems such as the Earth's crust and its oceans. The realm of geochemistry extends beyond the Earth, encompassing the entire Solar System, and has made important contributions to the understanding of a number of processes including mantle convection, the formation of planets and the origins of granite and basalt. It is an integrated field of chemistry and geology. History The term ''geochemistry'' was first used by the Swiss-German chemist Christian Friedrich Schönbein in 1838: "a comparative geochemistry ought to be launched, before geognosy can become geology, and before the mystery of the genesis of our planets and their inorganic matter may be revealed." However, for the rest of the century the more common term was "chemical geology", and there was little contact between geologists and chemists. Geochemistry emerged as a separate discipline after ...
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Franceville Basin
The Franceville Basin is a 1.6–2.1 billion year old sedimentary basis in Gabon. It contains unmetamorphosed sediments. It is notable for containing the Francevillian Biota, which is likely the oldest multicellular life known. A natural fission reactor formed there about 800-900 million years ago. Geology The Franceville Basin cover approximately 25,000 km2 and is made up of unmetamorphosed sediment derived mainly from eroded Mesoarchaean tonalite–trondhjemite–granodiorites. It is over a kilometer thick, with various sources claiming 2.5-4 kilometers as the maximum depth. Around 800-900 mya a natural fission reactor formed. The resulting fission by-products were held in place by a clay layer. See also *Francevillian Biota *Francevillian B Formation The Francevillian B Formation, also known as the Francevillian Formation or FB2 in scientific research, is a geologic formation of black shale provinces close to the town of Franceville, Gabon. The formation was depo ...
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Beltanelloides
''Beltanelliformis'' is a genus of discoid fossil from the Ediacaran period containing the two species B. brunsae and B. minutae, sometimes ascribed to the Ediacaran Biota. The chemical signature obtained from organically preserved specimens points to a cyanobacterial affinity (cf. ''Nostoc''). Depending on its preservation, it is sometimes referred to as ''Nemiana ''or ''Beltanelloides''. References

Ediacaran life Ediacaran Enigmatic prehistoric animal genera Aquatic animals Cyanobacteria {{ediacaran-stub ...
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Nemiana
''Beltanelliformis'' is a genus of discoid fossil from the Ediacaran period containing the two species B. brunsae and B. minutae, sometimes ascribed to the Ediacaran Biota. The chemical signature obtained from organically preserved specimens points to a cyanobacterial affinity (cf. ''Nostoc ''Nostoc'', also known as star jelly, troll’s butter, spit of moon, fallen star, witch's butter (not to be confused with the fungi commonly known as witches' butter), and witch’s jelly, is the most common genus of cyanobacteria found in var ...''). Depending on its preservation, it is sometimes referred to as ''Nemiana ''or ''Beltanelloides''. References Ediacaran life Ediacaran Enigmatic prehistoric animal genera Aquatic animals Cyanobacteria {{ediacaran-stub ...
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