Palaeoproterozoic
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The Paleoproterozoic Era (also spelled Palaeoproterozoic) is the first of the three sub-divisions ( eras) of the
Proterozoic The Proterozoic ( ) is the third of the four geologic eons of Earth's history, spanning the time interval from 2500 to 538.8 Mya, and is the longest eon of Earth's geologic time scale. It is preceded by the Archean and followed by the Phanerozo ...
eon, and also the longest era of the Earth's geological history, spanning from (2.5–1.6  Ga). It is further subdivided into four
geologic period The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronolo ...
s, namely the
Siderian The Siderian Period (; , meaning "iron") is the first geologic period in the Paleoproterozoic Era and lasted from Ma to Ma. Instead of being based on stratigraphy, these dates are defined chronometrically. The deposition of banded iron form ...
,
Rhyacian The Rhyacian Period (; , meaning "stream of lava") is the second geologic period in the Paleoproterozoic Era and lasted from Mya to Mya (million years ago). Instead of being based on stratigraphy, these dates are defined chronometrically. The ...
,
Orosirian The Orosirian Period (; , meaning "mountain range") is the third geologic period in the Paleoproterozoic Era and lasted from Mya to Mya (million years ago). Instead of being based on stratigraphy, these dates are defined chronometrically. T ...
and
Statherian The Statherian Period (; , meaning "stable, firm") is the final geologic period in the Paleoproterozoic Era and lasted from Mya to Mya (million years ago). Instead of being based on stratigraphy, these dates are defined chronometrically. Th ...
. Paleontological evidence suggests that the Earth's rotational rate ~1.8 billion years ago equated to 20-hour days, implying a total of ~450 days per year. It was during this era that the
continent A continent is any of several large geographical regions. Continents are generally identified by convention (norm), convention rather than any strict criteria. A continent could be a single large landmass, a part of a very large landmass, as ...
s first stabilized.


Atmosphere

The
Earth's atmosphere The atmosphere of Earth is composed of a layer of gas mixture that surrounds the Earth's planetary surface (both lands and oceans), known collectively as air, with variable quantities of suspended aerosols and particulates (which create weathe ...
was originally a weakly
reducing atmosphere A reducing atmosphere is an atmosphere in which oxidation is prevented by the absence of oxygen and other oxidizing gases or vapours, and which may contain actively reductant gases such as hydrogen, carbon monoxide, methane and hydrogen sulfide ...
consisting largely of
nitrogen Nitrogen is a chemical element; it has Symbol (chemistry), symbol N and atomic number 7. Nitrogen is a Nonmetal (chemistry), nonmetal and the lightest member of pnictogen, group 15 of the periodic table, often called the Pnictogen, pnictogens. ...
,
methane Methane ( , ) is a chemical compound with the chemical formula (one carbon atom bonded to four hydrogen atoms). It is a group-14 hydride, the simplest alkane, and the main constituent of natural gas. The abundance of methane on Earth makes ...
,
ammonia Ammonia is an inorganic chemical compound of nitrogen and hydrogen with the chemical formula, formula . A Binary compounds of hydrogen, stable binary hydride and the simplest pnictogen hydride, ammonia is a colourless gas with a distinctive pu ...
,
carbon dioxide Carbon dioxide is a chemical compound with the chemical formula . It is made up of molecules that each have one carbon atom covalent bond, covalently double bonded to two oxygen atoms. It is found in a gas state at room temperature and at norma ...
and
inert gas An inert gas is a gas that does not readily undergo chemical reactions with other chemical substances and therefore does not readily form chemical compounds. Though inert gases have a variety of applications, they are generally used to prevent u ...
es, in total comparable to Titan's atmosphere. When
oxygenic photosynthesis Photosynthesis ( ) is a system of biological processes by which photosynthetic organisms, such as most plants, algae, and cyanobacteria, convert light energy, typically from sunlight, into the chemical energy necessary to fuel their metaboli ...
evolved in
cyanobacteria Cyanobacteria ( ) are a group of autotrophic gram-negative bacteria that can obtain biological energy via oxygenic photosynthesis. The name "cyanobacteria" () refers to their bluish green (cyan) color, which forms the basis of cyanobacteri ...
during the
Mesoarchean The Mesoarchean ( , also spelled Mesoarchaean) is a geologic era in the Archean Eon, spanning , which contains the first evidence of modern-style plate subduction and expansion of microbial life. The era is defined chronometrically and is no ...
, the increasing amount of
byproduct A by-product or byproduct is a secondary product derived from a production process, manufacturing process or chemical reaction; it is not the primary product or service being produced. A by-product can be useful and marketable or it can be cons ...
dioxygen There are several known allotropes of oxygen. The most familiar is molecular oxygen (), present at significant levels in Earth's atmosphere and also known as dioxygen or triplet oxygen. Another is the highly reactive ozone (). Others are: * Ato ...
began to deplete the
reductant In chemistry, a reducing agent (also known as a reductant, reducer, or electron donor) is a chemical species that "donates" an electron to an (called the , , , or ). Examples of substances that are common reducing agents include hydrogen, carbon ...
s in the
ocean The ocean is the body of salt water that covers approximately 70.8% of Earth. The ocean is conventionally divided into large bodies of water, which are also referred to as ''oceans'' (the Pacific, Atlantic, Indian Ocean, Indian, Southern Ocean ...
,
land surface Terrain (), alternatively relief or topographical relief, is the dimension and shape of a given surface of land. In physical geography, terrain is the lay of the land. This is usually expressed in terms of the elevation, slope, and orientati ...
and the atmosphere. Eventually all surface reductants (particularly ferrous iron,
sulfur Sulfur ( American spelling and the preferred IUPAC name) or sulphur ( Commonwealth spelling) is a chemical element; it has symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms ...
and
atmospheric methane Atmospheric methane is the methane present in Earth's atmosphere. The concentration of atmospheric methane is increasing due to methane emissions, and is causing climate change. Methane is one of the most potent greenhouse gases. Methane's radiati ...
) were exhausted, and the atmospheric free oxygen levels soared permanently during the Siderian and Rhyacian periods in an aerochemical event called the
Great Oxidation Event The Great Oxidation Event (GOE) or Great Oxygenation Event, also called the Oxygen Catastrophe, Oxygen Revolution, Oxygen Crisis or Oxygen Holocaust, was a time interval during the Earth's Paleoproterozoic era when the Earth's atmosphere an ...
, which brought atmospheric oxygen from near none to up to 10% of the modern level.


Life

At the beginning of the preceding
Archean The Archean ( , also spelled Archaean or Archæan), in older sources sometimes called the Archaeozoic, is the second of the four geologic eons of Earth's history of Earth, history, preceded by the Hadean Eon and followed by the Proterozoic and t ...
eon, almost all existing lifeforms were single-cell
prokaryotic A prokaryote (; less commonly spelled procaryote) is a single-celled organism whose cell lacks a nucleus and other membrane-bound organelles. The word ''prokaryote'' comes from the Ancient Greek (), meaning 'before', and (), meaning 'nut' ...
anaerobic organism An anaerobic organism or anaerobe is any organism that does not require oxygen, molecular oxygen for growth. It may react negatively or even die if free oxygen is present. In contrast, an aerobic organism (aerobe) is an organism that requires an o ...
s whose
metabolism Metabolism (, from ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run cellular processes; the co ...
was based on a form of
cellular respiration Cellular respiration is the process of oxidizing biological fuels using an inorganic electron acceptor, such as oxygen, to drive production of adenosine triphosphate (ATP), which stores chemical energy in a biologically accessible form. Cell ...
that did not require oxygen, and
autotroph An autotroph is an organism that can convert Abiotic component, abiotic sources of energy into energy stored in organic compounds, which can be used by Heterotroph, other organisms. Autotrophs produce complex organic compounds (such as carbohy ...
s were either chemosynthetic or relied upon
anoxygenic photosynthesis Anoxygenic photosynthesis is a special form of photosynthesis used by some bacteria and archaea, which differs from the better known oxygenic photosynthesis in plants in the reductant used (e.g. hydrogen sulfide instead of water) and the byproduc ...
. After the Great Oxygenation Event, the then mainly
archaea Archaea ( ) is a Domain (biology), domain of organisms. Traditionally, Archaea only included its Prokaryote, prokaryotic members, but this has since been found to be paraphyletic, as eukaryotes are known to have evolved from archaea. Even thou ...
-dominated anaerobic
microbial mat A microbial mat is a multi-layered sheet or biofilm of microbial colonies, composed of mainly bacteria and/or archaea. Microbial mats grow at interfaces between different types of material, mostly on submerged or moist surfaces, but a few surviv ...
s were devastated as free oxygen is highly reactive and biologically toxic to cellular structures. This was compounded by a 300- million-year-long global icehouse event known as the Huronian glaciation — at least partly due to the depletion of atmospheric methane, a powerful
greenhouse gas Greenhouse gases (GHGs) are the gases in the atmosphere that raise the surface temperature of planets such as the Earth. Unlike other gases, greenhouse gases absorb the radiations that a planet emits, resulting in the greenhouse effect. T ...
— resulted in what is widely considered one of the first and most significant
mass extinction An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp fall in the diversity and abundance of multicellular organisms. It occ ...
s on Earth. The organisms that thrived after the extinction were mainly
aerobe An aerobic organism or aerobe is an organism that can survive and grow in an oxygenated environment. The ability to exhibit aerobic respiration may yield benefits to the aerobic organism, as aerobic respiration yields more energy than anaerobic ...
s that evolved bioactive antioxidants and eventually
aerobic respiration Cellular respiration is the process of oxidizing biological fuels using an inorganic electron acceptor, such as oxygen, to drive production of adenosine triphosphate (ATP), which stores chemical energy in a biologically accessible form. Cellu ...
, and surviving anaerobes were forced to live
symbiotic Symbiosis (Ancient Greek : living with, companionship < : together; and ''bíōsis'': living) is any type of a close and long-term biolo ...
ally alongside aerobes in hybrid colonies, which enabled the evolution of
mitochondria A mitochondrion () is an organelle found in the cells of most eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is us ...
in
eukaryotic organism The eukaryotes ( ) constitute the domain of Eukaryota or Eukarya, organisms whose cells have a membrane-bound nucleus. All animals, plants, fungi, seaweeds, and many unicellular organisms are eukaryotes. They constitute a major group of li ...
s. The Palaeoproterozoic represents the era from which the oldest cyanobacterial fossils, those of ''Eoentophysalis belcherensis'' from the Kasegalik Formation in the Belcher Islands of
Nunavut Nunavut is the largest and northernmost Provinces and territories of Canada#Territories, territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the ''Nunavut Act'' and the Nunavut Land Claims Agr ...
, are known. By 1.75 Ga, thylakoid-bearing cyanobacteria had evolved, as evidenced by fossils from the McDermott Formation of Australia. Many crown node eukaryotes (from which the modern-day eukaryotic lineages would have arisen) have been approximately dated to around the time of the Paleoproterozoic Era. While there is some debate as to the exact time at which eukaryotes evolved, current understanding places it somewhere in this era. Statherian
fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
from the Changcheng Group in
North China North China () is a list of regions of China, geographical region of the People's Republic of China, consisting of five province-level divisions of China, provincial-level administrative divisions, namely the direct-administered municipalities ...
provide evidence that eukaryotic life was already diverse by the late Palaeoproterozoic.


Geological events

During this era, the earliest global-scale continent-continent collision belts developed. The associated continent and mountain building events are represented by the 2.1–2.0 Ga Trans-Amazonian and Eburnean orogens in South America and West Africa; the ~2.0 Ga
Limpopo Belt The Limpopo Belt is located in South Africa and Zimbabwe, runs E-NE, and joins the Kaapvaal craton to the south with the Zimbabwe Craton to the north. The belt is of high-grade metamorphic rocks that have undergone a long cycle of metamorphism a ...
in southern Africa; the 1.9–1.8 Ga Trans-Hudson, Penokean, Taltson–Thelon, Wopmay, Ungava and Torngat orogens in North America, the 1.9–1.8 Ga Nagssugtoqidian Orogen in Greenland; the 1.9–1.8 Ga Kola–Karelia, Svecofennian, Volhyn-Central Russian, and Pachelma orogens in Baltica (Eastern Europe); the 1.9–1.8 Ga Akitkan Orogen in Siberia; the ~1.95 Ga Khondalite Belt; the ~1.85 Ga Trans-North China Orogen in North China; and the 1.8-1.6 Ga Yavapai and Mazatzal orogenies in southern North America. That pattern of collision belts supports the formation of a Proterozoic supercontinent named Columbia or Nuna. That continental collisions suddenly led to mountain building at large scale is interpreted as having resulted from increased biomass and carbon burial during and after the Great Oxidation Event: Subducted carbonaceous sediments are hypothesized to have lubricated compressive deformation and led to crustal thickening.
Felsic In geology, felsic is a grammatical modifier, modifier describing igneous rocks that are relatively rich in elements that form feldspar and quartz.Marshak, Stephen, 2009, ''Essentials of Geology,'' W. W. Norton & Company, 3rd ed. It is contrasted ...
volcanism in what is now northern Sweden led to the formation of the
Kiruna (; ; ; ) is the northernmost Stad (Sweden), city in Sweden, situated in the province of Lapland, Sweden, Lapland. It had 17,002 inhabitants in 2016 and is the seat of Kiruna Municipality (population: 23,167 in 2016) in Norrbotten County. The c ...
and Arvidsjaur porphyries. The lithospheric mantle of Patagonia's oldest blocks formed.


See also

* * * * * * , which immediately preceded the Paleoproterozoic


References


External links


EssayWeb Paleoproterozoic Era


New Scientist, #2746, 5 February 2010 by Nick Lane. Posits an earlier much longer snowball period, c2.4 - c2.0 Gya, triggered by the
Great Oxygenation Event The Great Oxidation Event (GOE) or Great Oxygenation Event, also called the Oxygen Catastrophe, Oxygen Revolution, Oxygen Crisis or Oxygen Holocaust, was a time interval during the Earth's Paleoproterozoic era when the Earth's atmosphere and ...
. * The information on eukaryotic lineage diversification was gathered from a
New York Times ''The New York Times'' (''NYT'') is an American daily newspaper based in New York City. ''The New York Times'' covers domestic, national, and international news, and publishes opinion pieces, investigative reports, and reviews. As one of ...
opinion blog by Olivia Judson. See the text here


Paleoproterozoic (chronostratigraphy scale)
{{Authority control Paleoproterozoic, Geological eras