Empidoidea
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Empidoidea
The Empidoidea are a large monophyletic superfamily of true flies, the sister taxon to the Muscomorpha (Cyclorrhapha). These two groups are sometimes united in the unranked taxon Eremoneura. There are some 10,000 known species within Empidoidea,Moulton & Wiegmann (2007) which are represented on all continents except Antarctica. They are known to have existed since the Jurassic period.Grimaldi & Engel (2005) Empidoidea has been subject to much debate regarding its phylogeny. Based on morphology alone, three major hypotheses had been proposed until 2007 and seemed to be consensus for some time—however, in 2018, Wahlberg & Johanson published the most current phylogeny supported by extensive genetic data, changing the relationship between quite a few families and tribes.Wahlberg & Johanson (2018) Description The majority of these insects are predatory, often with large compound eyes (sometimes covering almost the entire surface of the head), and tend to be associated with moist, t ...
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Empidoidea
The Empidoidea are a large monophyletic superfamily of true flies, the sister taxon to the Muscomorpha (Cyclorrhapha). These two groups are sometimes united in the unranked taxon Eremoneura. There are some 10,000 known species within Empidoidea,Moulton & Wiegmann (2007) which are represented on all continents except Antarctica. They are known to have existed since the Jurassic period.Grimaldi & Engel (2005) Empidoidea has been subject to much debate regarding its phylogeny. Based on morphology alone, three major hypotheses had been proposed until 2007 and seemed to be consensus for some time—however, in 2018, Wahlberg & Johanson published the most current phylogeny supported by extensive genetic data, changing the relationship between quite a few families and tribes.Wahlberg & Johanson (2018) Description The majority of these insects are predatory, often with large compound eyes (sometimes covering almost the entire surface of the head), and tend to be associated with moist, t ...
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Empididae
__NOTOC__ Empididae is a family of flies with over 3,000 described species occurring worldwide in all the biogeographic realms but the majority are found in the Holarctic. They are mainly predatory flies like most of their relatives in the Empidoidea, and exhibit a wide range of forms but are generally small to medium-sized, non-metallic and rather bristly. Common names for members of this family are dagger flies (referring to the sharp piercing mouthparts of some species) and balloon flies. The term "dance flies" is sometimes used for this family too, but the dance flies proper, formerly included herein, are now considered a separate family Hybotidae. Description For terms see Morphology of Diptera. Empididae are small to medium-sized flies, rarely large (1.0 to 15.0mm.). The body is slender, or elongated and rarely thickset. The colour ranges from yellow to black, and they may be pollinose or lustrous, but never have a metallic gloss. The head is often small and rounded with ...
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Dolichopodidae
Dolichopodidae, the long-legged flies, are a large, cosmopolitan family of true flies with more than 7,000 described species in about 230 genera. The genus ''Dolichopus'' is the most speciose, with some 600 species. Dolichopodidae generally are small flies with large, prominent eyes and a metallic cast to their appearance, though there is considerable variation among the species. Most have long legs, though some do not. In many species, the males have unusually large genitalia which are taxonomically useful in identifying species. Most adults are predatory on other small animals, though some may scavenge or act as kleptoparasites of spiders or other predators. An expanded concept of the family (Dolichopodidae ''sensu lato'') includes the subfamilies Parathalassiinae and Microphorinae. The latter of these was formerly placed in the Empididae, and was at one time considered a separate family (Microphoridae). However, some authors propose instead that Dolichopodidae ''s.l.'' shou ...
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Atelestidae
Atelestidae is a family of flies in the superfamily Empidoidea. The four genera were placed in a separate family in 1983; they were formerly either in Platypezidae (which are not even particularly closely related) or considered ''incertae sedis''. While they are doubtless the most basal of the living Empidoidea, the monophyly of the family is not fully proven. The genus '' Nemedina'' seems to represent a most ancient lineage among the entire superfamily, while '' Meghyperus'' is probably not monophyletic in its present delimitation, and it is liable to be split up eventually, with some species being placed elsewhere. In 2010, the genus '' Alavesia'', previously only known from Cretaceous fossils, was found alive in Namibia, subsequent species were also described from Brazil. Atelestidae has been shown to form the sister group to the remaining members of the Empidoidea superfamily. Subfamilies include Atelestinae and Nemedininae.Wahlberg & Johanson (2018) Description Atelestid ...
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Hybotidae
Hybotidae, the typical dance flies, are a family of true flies. They belong to the superfamily Empidoidea and were formerly included in the Empididae as a subfamily. Some, such as '' Tachydromia'', are predators that run around on the bark of trees in complex patterns, hence the common name. ''Tachydromia'' species are only about three millimeters long. Description Hybotidae share some similarities with the family Dolichopodidae, when looking at rotation of genitalia and wing characteristics. Male terminalia are rotated dextrally between 45° and 90°, excluding segment 7. Hybotidae wings always have a simple R4+5 vein, where the costa either ends near or at M1/M1+2, or near or at R4+5/R5. Furthermore, it can be distinguished from Dolichopodidae by the point of vein Rs, which it at a distance from the humeral crossvein (h) equal to or longer than the length of h. Systematics The Hybotidae clearly form a lineage quite distinct from the Empididae. Among the Empidoidea, they re ...
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Ragadidae
Ragadidae is a family of true flies in the superfamily Empidoidea. It was formerly considered a lower taxon, but was published as a new subfamily within Empididae in 2016.Sinclair (2016) Since then, it has been classified as the sister group to Empididae, and has been elevated to family level based on the genetic differences which separate it from Empididae.Wahlberg & Johanson (2018) Description Ragadidae are similar to Empididae and Atelestidae in the sense that their genitalia have symmetrical and straight terminalia. In their wings, the point of origin of the Rs vein is located at a distance from the humeral crossvein (h) as long as, or longer than, h. There is also a circumambient costa in the wing, which distinguishes Ragadidae from Atelestidae. Furthermore, the prosternum being separated from the proepisternum sets Ragadidae aside from Empididae—this is true for all except one genera (''Hydropeza'' ''spp''.), which can instead be characterized through its recurved labr ...
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Brachystomatinae
Brachystomatinae is a subfamily of flies belonging to the family Empididae. (2018): Molecular phylogenetics reveals novel relationships within Empidoidea (Diptera). ''Systematic Entomology'' 43(4): 619–636. Taxonomy Until 2006, Brachystomatinae was classified as a subfamily within Empididae, at which point a new phylogeny was proposed in which the lineage was raised to family rank, though other contemporaneous studies did not support this conclusion, (2007): The phylogenetic relationships of flies in the superfamily Empidoidea (Insecta: Diptera). '' Mol. Phylogenet. Evol.'' 43(3): 701-713. and in 2018 a new analysis indicated that the treatment of Brachystomatinae as a family rendered Empididae paraphyletic, and restored it to the rank of subfamily. Genera *'' Anomalempis'' Melander, 1928 *'' Apalocnemis'' Philippi, 1865 *'' Boreodromia'' Coquillett, 1903 *'' Brachystoma'' Meigen, 1822 *'' Ceratempis'' Melander, 1927 *'' Ceratomerus'' Philippi, 1865 *'' Ephydrempis'' Saigu ...
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Oreogetonidae
Oreogetonidae is a family of flies Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ... (insects in the order Diptera). There is at least one genus, '' Oreogeton'', and at least 30 described species in Oreogetonidae. References Further reading * * * * * * External links * Empidoidea Brachycera families {{Empidoidea-stub ...
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Muscomorpha
The Brachyceran infraorder Muscomorpha is a large and diverse group of flies, containing the bulk of the Brachycera, and, most of the known flies. It includes a number of the most familiar flies, such as the housefly, the fruit fly, and the blow fly. The antennae are short, usually three-segmented, with a dorsal arista. Their bodies are often highly setose, and the pattern of setae is often taxonomically important. The larvae of muscomorphs (in the sense the name is used here; see below) have reduced head capsules, and the pupae are formed inside the exoskeleton of the last larval instar; exit from this puparium is by a circular line of weakness, and this pupal type is called "cyclorrhaphous"; this feature gives this group of flies their traditional name, Cyclorrhapha. Classification The name Cyclorrhapha is used, in various modern classifications, to represent either a subgroup within the infraorder Muscomorpha, or simply a rankless group within the Brachycera. In either ca ...
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Homalocnemiidae
''Homalocnemis'' is a genus of flies which is placed in a family of its own, the Homalocnemiidae. There are about seven species in the genus found in the Afrotropical, Neotropical, and Australasian regions, suggestive of a Gondwanan origin. The genus was formerly considered a primitive empidoid and placed variously in the Hybotidae or in the empidid subfamily Brachystomatinae Brachystomatinae is a subfamily of flies belonging to the family Empididae. (2018): Molecular phylogenetics reveals novel relationships within Empidoidea (Diptera). ''Systematic Entomology'' 43(4): 619–636. Taxonomy Until 2006, Brachystomatina .... They are recognized by their wing venation which includes a long anal cell and a long basal segment of the antennal style. Species in the genus include: ;New Zealand * ''H. adelensis'' (Miller, 1913) * ''H. inexpleta'' Collin, 1928 * ''H. perspicua'' (Hutton, 1901) * ''H. maculipennis'' Malloch, 1932 ;Namibia * ''H. namibiensis'' Chvála, 1991 ;Chile * ''H. pr ...
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Diptera
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies and others, although only about 125,000 species have been described. Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great maneuverability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the l ...
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Eremoneura
Eremoneura is a clade of flies within the Brachycera that includes the Empidoidea and the Cyclorrhapha and is a sister of the Asilomorpha. They are thought to have evolved around the Mesozoic. The group includes fossils described in the genus '' Chimeromyia'' from 125 million year old amber which show both empidoid and cyclorrhaphan characters. The monotypic family Apystomyiidae has also been placed within the Eremoneura as a sister of the Cyclorrhapha Cyclorrhapha is an unranked taxon within the infraorder Muscomorpha. They are called "Cyclorrhapha" ('circular-seamed flies') with reference to the circular aperture through which the adult escapes the puparium. This is a circumscriptional name t .... References {{taxonbar, from=Q16634804 Muscomorpha ...
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