Euryzygomatomyinae
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Euryzygomatomyinae
Euryzygomatinae is a subfamily of rodents, proposed in 2017, and containing three extant genera of spiny Echimyidae: '' Clyomys'', '' Euryzygomatomys'', and ''Trinomys''. Members_of_this_echimyid_subfamily_all_share_an_origin_in_the_eastern_part_of_Brazil,_close_to_the_ e_...''. Members_of_this_echimyid_subfamily_all_share_an_origin_in_the_eastern_part_of_Brazil,_close_to_the_Atlantic_Forest. __Morpho-anatomy_ The_teeth_of_Euryzygomatomyines_are_characterized_by_several_features: *_elongate_lower_and_upper_incisor_roots_; *_five_lophids_on_the_lower_deciduous_premolars_4_; *_either_four_lophids_in_''Trinomys'',_or_three_lophids_in_''Clyomys''_and_''Euryzygomatomys'',_on_the_lower_molars_1_; *_well-connected_Molar_(tooth)#Lophodont.html" ;"title="Atlantic_Forest.html" ;"title="e ...''. Members of this echimyid subfamily all share an origin in the eastern part of Brazil, close to the Atlantic Forest">e ...''. Members of this echimyid subfamily all share an origin in the eastern p ...
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Echimyidae
Echimyidae is the family of neotropical spiny rats and their fossil relatives. This is the most species-rich family of hystricognath rodents. It is probably also the most ecologically diverse, with members ranging from fully arboreal to terrestrial to fossorial to semiaquatic habits. They presently exist mainly in South America; three members of the family also range into Central America, and the hutias are found in the Greater Antilles in the Caribbean. Species of the extinct subfamily Heteropsomyinae formerly lived on Cuba, Hispaniola, and Puerto Rico in the Antilles, probably until the arrival of Europeans. Characteristics In general form, most spiny rats resemble rats, although they are more closely related to guinea pigs and chinchillas. Most species have stiff, pointed hairs, or spines, that presumably serve for protection from predators. Many echimyids can break off their tails when attacked. This action may confuse predators long enough for the spiny rat to es ...
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Clyomys
''Clyomys'' is a South American rodent genus in the family Echimyidae. It contains two species, found in tropical savannas and grasslands from circa to elevation in central Brazil and eastern Paraguay. The term ''Clyomys'' derives from the two ancient greek words (, or ''clyo''), meaning "to listen, to prick up one's ears", and (), meaning "mouse, rat". The ''Clyomys '' species are as follows: * Broad-headed spiny rat (''Clyomys laticeps'') * † ''Clyomys riograndensis'' Phylogeny ''Clyomys'' is the sister genus to ''Euryzygomatomys''. Both taxa are closely related to the genus ''Trinomys''. In turn, these three genera — forming the clade of Euryzygomatomyinae — share phylogenetic affinities with a clade containing ''Carterodon Owl's spiny rat (''Carterodon sulcidens'') is a rodent species in the family Echimyidae found in Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South Ameri ...
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Euryzygomatomys
''Euryzygomatomys'' is a genus of South American rodents, commonly called guiaras, in the family Echimyidae. It contains two extant and one fossil species, found in Argentina, Brazil and Paraguay. They are as follows: * Brandt's guiara (''Euryzygomatomys guiara'') * Fischer's guiara (''Euryzygomatomys spinosus'') * †'' Euryzygomatomys hoffstetteri'' - Tarija Formation, BoliviaTarija
at Fossilworks.org


Etymology

The genus name ''Euryzygomatomys'' derives from the three words (, or ''eury''), meaning "wide, or which extends in width", or (), meaning "a part ...
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Trinomys
The Atlantic spiny rats are all found in the genus ''Trinomys''. They are a group of South American spiny-rats in the family Echimyidae. Extant species of ''Trinomys'' Based on Natureserve.InfoNatura: Animals and Ecosystems of Latin America [web application]. 2007. Version 5.0 . Arlington, Virginia (USA): NatureServe. Available: http://www.natureserve.org/infonatura. The species of spiny rats in the genus ''Trinomys'' are apparently all Brazilian. In particular many of them are endemic to the Atlantic Forest of southeastern Brazil. Not much is known for certain about their Range (biology), ranges however, which still are being investigated, with frequent extensions to the recorded distributions of several species in various ecological classes of forest and dry land. They do not however appear to occur at high altitudes, and several of their ranges appear to be parapatric. Most species of ''Trinomys'' are terrestrial and ambulatory, though ''Trinomys yonenagae'' is unusu ...
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Carterodon
Owl's spiny rat (''Carterodon sulcidens'') is a rodent species in the family Echimyidae found in Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area .... It is the only species in the genus ''Carterodon''. Owl's spiny rat has evolved characteristics such as a heightened ability to dig in open grasslands during times of environmental change.Bezerra, Alexandra & Marinho-Filho, Jader & Carmignotto, Ana. (2011). A review of the distribution, morphology, and habit of the Owl’s Spiny Rat Carterodon sulcidens (Lund, 1841) (Rodentia: Echimyidae). Zoological studies. 50. 566-576. Phylogeny The genus ''Carterodon'' is the sister group to the family Capromyidae (hutias). In turn, this clade shares evolutionary affinities with some genera of spiny rats belonging to the subfamily Euryzyg ...
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Atlantic Forest
The Atlantic Forest ( pt, Mata Atlântica) is a South American forest that extends along the Atlantic coast of Brazil from Rio Grande do Norte state in the northeast to Rio Grande do Sul state in the south and inland as far as Paraguay and the Misiones Province of Argentina, where the region is known as Selva Misionera. The Atlantic Forest has ecoregions within the following biome categories: seasonal moist and dry broad-leaf tropical forests, tropical and subtropical grasslands, savannas, and shrublands, and mangrove forests. The Atlantic Forest is characterized by a high biodiversity and endemism. It was the first environment that the Portuguese colonists encountered over 500 years ago, when it was thought to have had an area of , and stretching an unknown distance inland, making it, back then, the second largest rainforest on the planet, only behind the Amazon rainforest. Over 85% of the original area has been deforested, threatening many plant and animal species with ...
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Molar (tooth)
The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone tooth", from ''mola'', millstone and ''dens'', tooth. Molars show a great deal of diversity in size and shape across mammal groups. The third molar of humans is sometimes vestigial. Human anatomy In humans, the molar teeth have either four or five cusps. Adult humans have 12 molars, in four groups of three at the back of the mouth. The third, rearmost molar in each group is called a wisdom tooth. It is the last tooth to appear, breaking through the front of the gum at about the age of 20, although this varies from individual to individual. Race can also affect the age at which this occurs, with statistical variations between groups. In some cases, it may not even erupt at all. The human mouth contains upper (maxillary) and lower (mandib ...
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Zygomatic Arch
In anatomy, the zygomatic arch, or cheek bone, is a part of the skull formed by the zygomatic process of the temporal bone (a bone extending forward from the side of the skull, over the opening of the ear) and the temporal process of the zygomatic bone (the side of the cheekbone), the two being united by an oblique suture (the zygomaticotemporal suture); the tendon of the temporal muscle passes medial to (i.e. through the middle of) the arch, to gain insertion into the coronoid process of the mandible (jawbone). The jugal point is the point at the anterior (towards face) end of the upper border of the zygomatic arch where the masseteric and maxillary edges meet at an angle, and where it meets the process of the zygomatic bone. The arch is typical of '' Synapsida'' (“fused arch”), a clade of amniotes that includes mammals and their extinct relatives, such as ''Moschops'' and '' Dimetrodon''. Structure The zygomatic process of the temporal arises by two roots: * an ''anter ...
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Jugal Bone
The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy The jugal bone is located on either side of the skull in the circumorbital region. It is the origin of several masticatory muscles in the skull. The jugal and lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones. During development, the jugal bone originates from dermal bone. In dinosaurs This bone is considered key in the determination of general traits in cases in which the entire skull has not been found intact (for instance, as with dinosaurs in paleontology). In some dinosaur genera the jugal also forms part of the lower margin of either the antorbital fenestra or the infratemporal fenestra, or both. Most commonly, this bone articu ...
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Leucine
Leucine (symbol Leu or L) is an essential amino acid that is used in the biosynthesis of proteins. Leucine is an α-amino acid, meaning it contains an α-amino group (which is in the protonated −NH3+ form under biological conditions), an α- carboxylic acid group (which is in the deprotonated −COO− form under biological conditions), and a side chain isobutyl group, making it a non-polar aliphatic amino acid. It is essential in humans, meaning the body cannot synthesize it: it must be obtained from the diet. Human dietary sources are foods that contain protein, such as meats, dairy products, soy products, and beans and other legumes. It is encoded by the codons UUA, UUG, CUU, CUC, CUA, and CUG. Like valine and isoleucine, leucine is a branched-chain amino acid. The primary metabolic end products of leucine metabolism are acetyl-CoA and acetoacetate; consequently, it is one of the two exclusively ketogenic amino acids, with lysine being the other. It is the most import ...
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Proline
Proline (symbol Pro or P) is an organic acid classed as a proteinogenic amino acid (used in the biosynthesis of proteins), although it does not contain the amino group but is rather a secondary amine. The secondary amine nitrogen is in the protonated form (NH2+) under biological conditions, while the carboxyl group is in the deprotonated −COO− form. The "side chain" from the α carbon connects to the nitrogen forming a pyrrolidine loop, classifying it as a aliphatic amino acid. It is non-essential in humans, meaning the body can synthesize it from the non-essential amino acid L-glutamate. It is encoded by all the codons starting with CC (CCU, CCC, CCA, and CCG). Proline is the only proteinogenic secondary amino acid which is a secondary amine, as the nitrogen atom is attached both to the α-carbon and to a chain of three carbons that together form a five-membered ring. History and etymology Proline was first isolated in 1900 by Richard Willstätter who obtained the amino ...
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Fossorial
A fossorial () animal is one adapted to digging which lives primarily, but not solely, underground. Some examples are badgers, naked mole-rats, clams, meerkats, and mole salamanders, as well as many beetles, wasps, and bees. Prehistoric evidence The physical adaptation of fossoriality is widely accepted as being widespread among many prehistoric phyla and taxa, such as bacteria and early eukaryotes. Furthermore, fossoriality has evolved independently multiple times, even within a single family. Fossorial animals appeared simultaneously with the colonization of land by arthropods in the late Ordovician period (over 440 million years ago). Other notable early burrowers include ''Eocaecilia'' and possibly ''Dinilysia''. The oldest example of burrowing in synapsids, the lineage which includes modern mammals and their ancestors, is a cynodont, ''Thrinaxodon liorhinus'', found in the Karoo of South Africa, estimated to be 251 million years old. Evidence shows that this ...
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