Empididae
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Empididae
__NOTOC__ Empididae is a family of flies with over 3,000 described species occurring worldwide in all the biogeographic realms but the majority are found in the Holarctic. They are mainly predatory flies like most of their relatives in the Empidoidea, and exhibit a wide range of forms but are generally small to medium-sized, non-metallic and rather bristly. Common names for members of this family are dagger flies (referring to the sharp piercing mouthparts of some species) and balloon flies. The term "dance flies" is sometimes used for this family too, but the dance flies proper, formerly included herein, are now considered a separate family Hybotidae. Description For terms see Morphology of Diptera. Empididae are small to medium-sized flies, rarely large (1.0 to 15.0mm.). The body is slender, or elongated and rarely thickset. The colour ranges from yellow to black, and they may be pollinose or lustrous, but never have a metallic gloss. The head is often small and rounded with ...
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Brachystomatinae
Brachystomatinae is a subfamily of flies belonging to the family Empididae. (2018): Molecular phylogenetics reveals novel relationships within Empidoidea (Diptera). ''Systematic Entomology'' 43(4): 619–636. Taxonomy Until 2006, Brachystomatinae was classified as a subfamily within Empididae, at which point a new phylogeny was proposed in which the lineage was raised to family rank, though other contemporaneous studies did not support this conclusion, (2007): The phylogenetic relationships of flies in the superfamily Empidoidea (Insecta: Diptera). '' Mol. Phylogenet. Evol.'' 43(3): 701-713. and in 2018 a new analysis indicated that the treatment of Brachystomatinae as a family rendered Empididae paraphyletic, and restored it to the rank of subfamily. Genera *'' Anomalempis'' Melander, 1928 *'' Apalocnemis'' Philippi, 1865 *'' Boreodromia'' Coquillett, 1903 *'' Brachystoma'' Meigen, 1822 *'' Ceratempis'' Melander, 1927 *'' Ceratomerus'' Philippi, 1865 *'' Ephydrempis'' Saigu ...
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Empidinae
Empidinae are a subfamily of empidoid flies. They are mainly predatory flies like most of their relatives, and generally small to medium-sized. Most species are flower visitors and they can be effective pollinators. Genera *'' Afroempis'' Smith, 1969 *'' Allochrotus'' Collin, 1933 *'' Amictoides'' Bezzi, 1909 *'' Aplomera'' Macquart, 1838 *'' Atrichopleura'' Bezzi, 1909 *'' Bandella'' Bickel, 2002 *'' Clinorhampha'' Collin, 1933 *'' Cunomyia'' Bickel, 1998 *'' Deuteragonista'' Philippi, 1865 *'' Empidadelpha'' Collin, 1928 *''Empis'' Linnaeus, 1758 *'' Gynatoma'' Collin, 1928 *'' Hilara'' Meigen, 1822 *'' Hilarempis'' Bezzi, 1905 *'' Hilarigona'' Collin, 1933 *'' Hybomyia'' Plant, 1995 *'' Hystrichonotus'' Collin, 1933 *'' Lamprempis'' Wheeler & Melander, 1901 *'' Macrostomus'' Wiedemann, 1817 *'' Opeatocerata'' Melander, 1928 *'' Pasitrichotus'' Collin, 1933 *'' Porphyrochroa'' Melander, 1928 *'' Rhamphella'' Malloch, 1930 *''Rhamphomyia'' Meigen, 1822 *'' Sphi ...
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Hemerodromiinae
Hemerodromiinae are a worldwide group of predatory flies with raptorial forelegs.Plant, A.R. 2011: Hemerodromiinae (Diptera: Empididae): a tentative phylogeny and biogeographical discussion. ''Systematic entomology'', 36(1): 83-103. Genera *'' Afrodromia'' Smith, 1969 *'' Antipodromia'' Plant, 2011 *'' Chelifera'' Macquart, 1823 *'' Chelipoda'' Macquart, 1823 *'' Chelipodozus'' Collin, 1933 *'' Cladodromia'' Bezzi, 1905 *'' Colabris'' Melander, 1928 *'' Doliodromia'' Collin, 1928 *'' Drymodromia'' Bezzi, 1914 *'' Dryodromya'' Rondani, 1856 *'' Hemerodromia'' Meigen, 1822 *'' Metachela'' Coquillett, 1903 *'' Monodromia'' Collin, 1928 *'' Neoplasta'' Coquillett, 1895 *'' Phyllodromia'' Zetterstedt Johan Wilhelm Zetterstedt (20 May 1785 – 23 December 1874) was a Sweden, Swedish Naturalism (philosophy) , naturalist who worked mainly on Diptera and Hymenoptera. Biography Zetterstedt studied at the Lund University, University of Lund, where ..., 1837 *'' Ptilophyllodro ...
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Trichopezinae
Trichopezinae are a subfamily of empidoid flies. They are mainly predatory flies like most of their relatives, and generally small to medium-sized, long-legged and large-eyed. Previously, they were included in the Clinocerinae or the Hemerodromiinae. In some more recent treatments, the Brachystomatinae are considered a distinct family, and the Trichopezinae are placed therein. However, it is more likely that the Brachystomatinae are part of the Empididae, and that the Trichopezinae represent a separate lineage in the same family. The monophyly of the Trichopezinae versus its closest relatives is not firmly established; it seems likely that some changes will be necessary to make this subfamily a natural group. Selected genera *''Apalocnemis'' Philippi, 1865 *'' Boreodromia'' Coquillett, 1903 *'' Ceratempis'' Melander, 1927 *''Ephydrempis'' Saigusa, 1986 *''Gloma'' Meigen, 1822 *'' Heleodromia'' Haliday, 1833 *'' Heterophlebus'' Philippi Philippi (; grc-gre, Φίλιππο ...
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Ragadidae
Ragadidae is a family of true flies in the superfamily Empidoidea. It was formerly considered a lower taxon, but was published as a new subfamily within Empididae in 2016.Sinclair (2016) Since then, it has been classified as the sister group to Empididae, and has been elevated to family level based on the genetic differences which separate it from Empididae.Wahlberg & Johanson (2018) Description Ragadidae are similar to Empididae and Atelestidae in the sense that their genitalia have symmetrical and straight terminalia. In their wings, the point of origin of the Rs vein is located at a distance from the humeral crossvein (h) as long as, or longer than, h. There is also a circumambient costa in the wing, which distinguishes Ragadidae from Atelestidae. Furthermore, the prosternum being separated from the proepisternum sets Ragadidae aside from Empididae—this is true for all except one genera (''Hydropeza'' ''spp''.), which can instead be characterized through its recurved labr ...
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Clinocerinae
Clinocerinae is a subfamily of flies belonging to the family Empididae. Genera *'' Afroclinocera'' Sinclair, 1999 *'' Asymphyloptera'' Collin, 1933 *'' Bergenstammia'' Mik, 1881 *'' Clinocera'' Meigen, 1803 *'' Dolichocephala'' Macquart, 1823 *'' Hypenella'' Collin, 1941 *'' Kowarzia'' Mik, 1881 *'' Oreothalia'' Melander, 1902 *'' Phaeobalia'' Mik, 1881 *'' Proagomyia'' Collin, 1933 *'' Proclinopyga'' Melander, 1928 *'' Rhyacodromia'' Saigusa, 1986 *'' Roederiodes'' Coquillett, 1901 *'' Trichoclinocera'' Collin, 1941 *'' Wiedemannia'' Zetterstedt Johan Wilhelm Zetterstedt (20 May 1785 – 23 December 1874) was a Sweden, Swedish Naturalism (philosophy) , naturalist who worked mainly on Diptera and Hymenoptera. Biography Zetterstedt studied at the Lund University, University of Lund, where ..., 1838 References Empididae Articles containing video clips Asilomorpha subfamilies {{empidoidea-stub ...
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Hybotidae
Hybotidae, the typical dance flies, are a family of true flies. They belong to the superfamily Empidoidea and were formerly included in the Empididae as a subfamily. Some, such as '' Tachydromia'', are predators that run around on the bark of trees in complex patterns, hence the common name. ''Tachydromia'' species are only about three millimeters long. Description Hybotidae share some similarities with the family Dolichopodidae, when looking at rotation of genitalia and wing characteristics. Male terminalia are rotated dextrally between 45° and 90°, excluding segment 7. Hybotidae wings always have a simple R4+5 vein, where the costa either ends near or at M1/M1+2, or near or at R4+5/R5. Furthermore, it can be distinguished from Dolichopodidae by the point of vein Rs, which it at a distance from the humeral crossvein (h) equal to or longer than the length of h. Systematics The Hybotidae clearly form a lineage quite distinct from the Empididae. Among the Empidoidea, they re ...
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Empis Livida
''Empis livida'' is a species of dance fly, in the fly family Empididae. It is included in the subgenus ''Kritempis'' of the genus ''Empis''. Males range from , females . The male's abdomen is brownish, and his wings appear faintly brown and clouded. The female's abdomen is gray and her wings are clear. ''E. livida'' lives in hedgerows, feeding on the nectar of several species of Rosaceae, several species of Asteraceae, and ''Heracleum sphondylium ''Heracleum sphondylium'', commonly known as hogweed, common hogweed or cow parsnip, is a herbaceous perennial or biennial plant, in the umbelliferous family Apiaceae that includes fennel, cow parsley, ground elder and giant hogweed. It is nat ...'' nectar; they also feed on other insects. They live all across temperate and Northern Europe, the only species with such a wide distribution. ''E. livida'' larvae are also carnivorous and live in damp soil and leaf litter. Adults fly in between April to July. References Empis Asi ...
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Empidoidea
The Empidoidea are a large monophyletic superfamily of true flies, the sister taxon to the Muscomorpha (Cyclorrhapha). These two groups are sometimes united in the unranked taxon Eremoneura. There are some 10,000 known species within Empidoidea,Moulton & Wiegmann (2007) which are represented on all continents except Antarctica. They are known to have existed since the Jurassic period.Grimaldi & Engel (2005) Empidoidea has been subject to much debate regarding its phylogeny. Based on morphology alone, three major hypotheses had been proposed until 2007 and seemed to be consensus for some time—however, in 2018, Wahlberg & Johanson published the most current phylogeny supported by extensive genetic data, changing the relationship between quite a few families and tribes.Wahlberg & Johanson (2018) Description The majority of these insects are predatory, often with large compound eyes (sometimes covering almost the entire surface of the head), and tend to be associated with moist, t ...
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Dolichopodidae
Dolichopodidae, the long-legged flies, are a large, cosmopolitan family of true flies with more than 7,000 described species in about 230 genera. The genus ''Dolichopus'' is the most speciose, with some 600 species. Dolichopodidae generally are small flies with large, prominent eyes and a metallic cast to their appearance, though there is considerable variation among the species. Most have long legs, though some do not. In many species, the males have unusually large genitalia which are taxonomically useful in identifying species. Most adults are predatory on other small animals, though some may scavenge or act as kleptoparasites of spiders or other predators. An expanded concept of the family (Dolichopodidae ''sensu lato'') includes the subfamilies Parathalassiinae and Microphorinae. The latter of these was formerly placed in the Empididae, and was at one time considered a separate family (Microphoridae). However, some authors propose instead that Dolichopodidae ''s.l.'' shou ...
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Atelestidae
Atelestidae is a family of flies in the superfamily Empidoidea. The four genera were placed in a separate family in 1983; they were formerly either in Platypezidae (which are not even particularly closely related) or considered ''incertae sedis''. While they are doubtless the most basal of the living Empidoidea, the monophyly of the family is not fully proven. The genus '' Nemedina'' seems to represent a most ancient lineage among the entire superfamily, while '' Meghyperus'' is probably not monophyletic in its present delimitation, and it is liable to be split up eventually, with some species being placed elsewhere. In 2010, the genus '' Alavesia'', previously only known from Cretaceous fossils, was found alive in Namibia, subsequent species were also described from Brazil. Atelestidae has been shown to form the sister group to the remaining members of the Empidoidea superfamily. Subfamilies include Atelestinae and Nemedininae.Wahlberg & Johanson (2018) Description Atelestid ...
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Diptera
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies and others, although only about 125,000 species have been described. Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great maneuverability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the l ...
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