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Cygnus Columbianus
The tundra swan (''Cygnus columbianus'') is a small swan of the Holarctic. The two taxa within it are usually regarded as conspecific, but are also sometimes split into two species: Bewick's swan (''Cygnus bewickii'') of the Palaearctic and the whistling swan (''C. columbianus'') proper of the Nearctic. Birds from eastern Russia (roughly east of the Taymyr Peninsula) are sometimes separated as the subspecies ''C. c. jankowskii'', but this is not widely accepted as distinct, with most authors including them in ''C. c. bewickii''. Tundra swans are sometimes separated in the subgenus ''Olor'' together with the other Arctic swan species. Bewick's swan was named in 1830 by William Yarrell after the engraver Thomas Bewick, who specialised in illustrations of birds and animals. ''Cygnus'' is the Latin for "swan", and '' columbianus'' comes from the Columbia River, the type locality. Description ''C. columbianus'' is the smallest of the Holarctic swans, at in length, in wingspan a ...
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Gloucestershire
Gloucestershire ( abbreviated Glos) is a county in South West England. The county comprises part of the Cotswold Hills, part of the flat fertile valley of the River Severn and the entire Forest of Dean. The county town is the city of Gloucester and other principal towns and villages include Cheltenham, Cirencester, Kingswood, Bradley Stoke, Stroud, Thornbury, Yate, Tewkesbury, Bishop's Cleeve, Churchdown, Brockworth, Winchcombe, Dursley, Cam, Berkeley, Wotton-under-Edge, Tetbury, Moreton-in-Marsh, Fairford, Lechlade, Northleach, Stow-on-the-Wold, Chipping Campden, Bourton-on-the-Water, Stonehouse, Nailsworth, Minchinhampton, Painswick, Winterbourne, Frampton Cotterell, Coleford, Cinderford, Lydney and Rodborough and Cainscross that are within Stroud's urban area. Gloucestershire borders Herefordshire to the north-west, Worcestershire to the north, Warwickshire to the north-east, Oxfordshire to the east, Wiltshire to the south, Bristol and Somerset ...
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Hybrid (biology)
In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Hybrids are not always intermediates between their parents (such as in blending inheritance), but can show hybrid vigor, sometimes growing larger or taller than either parent. The concept of a hybrid is interpreted differently in animal and plant breeding, where there is interest in the individual parentage. In genetics, attention is focused on the numbers of chromosomes. In taxonomy, a key question is how closely related the parent species are. Species are reproductively isolated by strong barriers to hybridisation, which include genetic and morphological differences, differing times of fertility, mating behaviors and cues, and physiological rejection of sperm cells or the developing embryo. Some act before fertilization and others after it. Similar barriers exist in plants, with differences in flowering tim ...
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Beringia
Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip of the Kamchatka Peninsula. It includes the Chukchi Sea, the Bering Sea, the Bering Strait, the Chukchi and Kamchatka Peninsulas in Russia as well as Alaska in the United States and the Yukon in Canada. The area includes land lying on the North American Plate and Siberian land east of the Chersky Range. At certain times in prehistory, it formed a land bridge that was up to wide at its greatest extent and which covered an area as large as British Columbia and Alberta together, totaling approximately . Today, the only land that is visible from the central part of the Bering land bridge are the Diomede Islands, the Pribilof Islands of St. Paul and St. George, St. Lawrence Island, St. Matthew Island, and King Island. The term ''Beringi ...
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Gene Flow
In population genetics, gene flow (also known as gene migration or geneflow and allele flow) is the transfer of genetic material from one population to another. If the rate of gene flow is high enough, then two populations will have equivalent allele frequencies and therefore can be considered a single effective population. It has been shown that it takes only "one migrant per generation" to prevent populations from diverging due to drift. Populations can diverge due to selection even when they are exchanging alleles, if the selection pressure is strong enough. Gene flow is an important mechanism for transferring genetic diversity among populations. Migrants change the distribution of genetic diversity among populations, by modifying allele frequencies (the proportion of members carrying a particular variant of a gene). High rates of gene flow can reduce the genetic differentiation between the two groups, increasing homogeneity. For this reason, gene flow has been thought to con ...
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Beak
The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food, courtship, and feeding young. The terms ''beak'' and ''rostrum'' are also used to refer to a similar mouth part in some ornithischians, pterosaurs, cetaceans, dicynodonts, anuran tadpoles, monotremes (i.e. echidnas and platypuses, which have a beak-like structure), sirens, pufferfish, billfishes and cephalopods. Although beaks vary significantly in size, shape, color and texture, they share a similar underlying structure. Two bony projections – the upper and lower mandibles – are covered with a thin keratinized layer of epidermis known as the rhamphotheca. In most species, two holes called ''nares'' lead to the respiratory system. Etymology Although the word "beak" was, in the past, generally restricted to the sharpened bills o ...
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Whooper Swan
The whooper swan ( /ˈhuːpə(ɹ) swɒn/) (''Cygnus cygnus''), also known as the common swan, pronounced ''hooper swan'', is a large northern hemisphere swan. It is the Eurasian counterpart of the North American trumpeter swan, and the type species for the genus '' Cygnus''. Taxonomy Francis Willughby and John Ray's ''Ornithology'' of 1676 referred to this swan as "the Elk, Hooper, or wild Swan". It was one of the many bird species originally described by Carl Linnaeus in the 1758 10th edition of his ''Systema Naturae'', where it was given the binomial name of ''Anas cygnus''. The species name is from ''cygnus'', the Latin for "swan". Description The whooper swan is similar in appearance to Bewick's swan. It is larger, however, at a length of and a wingspan of . The weight is typically in the range of , with an average of for males and for females. The verified record mass was for a wintering male from Denmark. It is considered to be amongst the heaviest flying birds. Amon ...
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Parapatric Speciation
In parapatric speciation, two subpopulations of a species evolve reproductive isolation from one another while continuing to exchange genes. This mode of speciation has three distinguishing characteristics: 1) mating occurs non-randomly, 2) gene flow occurs unequally, and 3) populations exist in either continuous or discontinuous geographic ranges. This distribution pattern may be the result of unequal dispersal, incomplete geographical barriers, or divergent expressions of behavior, among other things. Parapatric speciation predicts that hybrid zones will often exist at the junction between the two populations. In biogeography, the terms parapatric and parapatry are often used to describe the relationship between organisms whose ranges do not significantly overlap but are immediately adjacent to each other; they do not occur together except in a narrow contact zone. Parapatry is a geographical distribution opposed to sympatry (same area) and allopatry or peripatry (two simila ...
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Tarsus (skeleton)
In the human body, the tarsus is a cluster of seven articulating bones in each foot situated between the lower end of the tibia and the fibula of the lower leg and the metatarsus. It is made up of the midfoot (Cuboid bone, cuboid, medial, intermediate, and lateral cuneiform bone, cuneiform, and navicular) and hindfoot (Talus bone, talus and calcaneus). The tarsus articulates with the bones of the metatarsus, which in turn articulate with the proximal phalanges of the toes. The joint between the tibia and fibula above and the tarsus below is referred to as the ankle, ankle joint proper. In humans the largest bone in the tarsus is the calcaneus, which is the weight-bearing bone within the heel of the foot. Human anatomy Bones The talus bone or ankle bone is connected superiorly to the two bones of the lower leg, the tibia and fibula, to form the ankle, ankle joint or talocrural joint; inferiorly, at the subtalar joint, to the calcaneus or heel bone. Together, the talus and ...
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Cline (biology)
In biology, a cline (from the Greek κλίνειν ''klinein'', meaning "to lean") is a measurable gradient in a single character (or biological trait) of a species across its geographical range. First coined by Julian Huxley in 1938, the "character" of the cline referred to is usually genetic (e.g. allele frequency, blood type), or phenotypic (e.g. body size, skin pigmentation). Clines can show smooth, continuous gradation in a character, or they may show more abrupt changes in the trait from one geographic region to the next. A cline refers to a spatial gradient in a specific, singular trait, rather than a collection of traits; a single population can therefore have as many clines as it has traits, at least in principle. Additionally, Huxley recognised that these multiple independent clines may not act in concordance with each other. For example, it has been observed that in Australia, birds generally become smaller the further towards the north of the country they are found ...
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Down Feather
The down of birds is a layer of fine feathers found under the tougher exterior feathers. Very young birds are clad only in down. Powder down is a specialized type of down found only in a few groups of birds. Down is a fine thermal insulator and padding, used in goods such as jackets, bedding (duvets and featherbeds), pillows and sleeping bags. The discovery of feathers trapped in ancient amber suggests that some species of dinosaur may have possessed down-like feathers. Description and etymology The word ''down'' comes from the Old Norse word ''dúnn'', which had the same meaning as its modern equivalent. The down feather is considered to be the most "straightforward" of all feather types. It has a short or vestigial rachis (shaft), few barbs, and barbules that lack hooks. There are three types of down: natal down, body down and powder down. Natal down is the layer of down feathers that cover most birds at some point in their early development. Precocial nestlings are alre ...
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Moulting
In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer layer or covering), either at specific times of the year, or at specific points in its life cycle. In medieval times it was also known as "mewing" (from the French verb "muer", to moult), a term that lives on in the name of Britain's Royal Mews where the King's hawks used to be kept during moulting time before becoming horse stables after Tudor times. Moulting can involve shedding the Epidermis (skin), epidermis (skin), pelage (hair, feathers, fur, wool), or other external layer. In some groups, other body parts may be shed, for example, the entire exoskeleton in arthropods, including the wings in some insects. Examples In birds In birds, moulting is the periodic replacement of feathers by shedding old feathers while producing new o ...
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