Chaoyangopterids
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Chaoyangopterids
Chaoyangopteridae (or chaoyangopterids) is a family of pterosaurs within the larger group Azhdarchoidea. Chaoyangopterids lived mostly during the Early Cretaceous period, though possible members, ''Microtuban'', '' Xericeps'' and ''Argentinadraco'', may extend the fossil range to the Late Cretaceous. History The clade Chaoyangopteridae was first defined in 2008 by Lü Junchang and David Unwin as: "''Chaoyangopterus'', ''Shenzhoupterus'', their most recent common ancestor and all taxa more closely related to this clade than to '' Tapejara'', ''Tupuxuara'' or ''Quetzalcoatlus''". Based on neck and limb proportions, it has been suggested they occupied a similar ecological niche to that of azhdarchid pterosaurs, though it is possible they were more specialized as several genera occur in Liaoning, while azhdarchids usually occur by one genus in a specific location. Description Chaoyangopterids are distinguished from other pterosaurs by several traits of the nasoantorbital fenestra, a l ...
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Chaoyangopterus
''Chaoyangopterus'' is a genus of chaoyangopterid pterosaur known from a partial skeleton found in Liaoning, China. ''Chaoyangopterus'' was found in rocks dating back to the Aptian-age Lower Cretaceous Jiufotang Formation of Dapingfang, Chaoyang. Etymology ''Chaoyangopterus'' was named and described in 2003 by Wang Xiaolin and Zhou Zhonghe. The type species is ''Chaoyangopterus zhangi''. The genus name is derived from Chaoyang and a Latinized Greek ''pteron'', "wing". The specific name honors journalist Zhang Wanlian for his efforts in protecting fossil sites. Description ''Chaoyangopterus'' is based on holotype IVPP V13397, which includes the front of the skull, the lower jaws, the neck vertebrae, the shoulder and pelvic girdles, and the limbs. The skull is about long and toothless, and its wingspan is estimated to have been around . Wang and Zhou concluded that it compared most closely to ''Nyctosaurus'' and classified it as a nyctosaurid, although they found that i ...
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Chaoyangopterus Zhangi
''Chaoyangopterus'' is a genus of chaoyangopterid pterosaur known from a partial skeleton found in Liaoning, China. ''Chaoyangopterus'' was found in rocks dating back to the Aptian-age Lower Cretaceous Jiufotang Formation of Dapingfang, Chaoyang. Etymology ''Chaoyangopterus'' was named and described in 2003 by Wang Xiaolin and Zhou Zhonghe. The type species is ''Chaoyangopterus zhangi''. The genus name is derived from Chaoyang and a Latinized Greek ''pteron'', "wing". The specific name honors journalist Zhang Wanlian for his efforts in protecting fossil sites. Description ''Chaoyangopterus'' is based on holotype IVPP V13397, which includes the front of the skull, the lower jaws, the neck vertebrae, the shoulder and pelvic girdles, and the limbs. The skull is about long and toothless, and its wingspan is estimated to have been around . Wang and Zhou concluded that it compared most closely to ''Nyctosaurus'' and classified it as a nyctosaurid, although they found that i ...
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Microtuban
''Microtuban'' is an extinct genus of azhdarchoid pterosaur known from the Late Cretaceous of northern Lebanon. Discovery ''Microtuban'' is known only from a partially preserved skeleton lacking the skull, holotype SMNK PAL 6595. It was acquired from a local fossil dealer by the ''Staatliches Museum für Naturkunde Karlsruhe'' and was, as can be deduced from the qualities of the stone matrix, collected in the locality of Hjoûla in a marine layer of the Sannine Formation, dating to the early Cenomanian stage of the Late Cretaceous, about 99.6–96 million years ago. The fossil dealer however, indicated it came from a quarry at nearby Hâqel. Regardless, it is only the second pterosaur fossil found in Lebanon and a rare example of a pterosaur from the African continental plate, to which the area in Cretaceous times belonged. In this period the site was located hundreds of miles from the mainland coastline. The specimen consists of the last cervical and first dorsal vertebrae, a ...
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Shenzhoupterus
''Shenzhoupterus'' is a genus of chaoyangopterid pterosaur from the Jiufotang Formation of modern-day Liaoning, China. Fossil remains of ''Shenzhoupterus'' date back to the Early Cretaceous period, approximately 120 million years ago. Etymology ''Shenzhoupterus'' was named in 2008 by Lü Junchang, David Unwin, Xu Li and Zhang Xingliao. The type species is ''Shenzoupterus chaoyangensis''. The genus name is derived from ''Shenzhou'', an old name for China, "the divine land", and a Latinized Greek ''pteron'', "wing". The specific name refers to Chaoyang. Description ''Shenzhoupterus'' is based on holotype HGM 41HIII-305A (Henan Geological Museum at Zhengzhou), the articulated skull and skeleton of a single individual, with a wingspan of . ''Shenzhoupterus'' lacked teeth, and had a crest on its skull that arched over the eyes and terminated in a small point toward the back of the head. The nasoantorbital fenestra (an opening incorporating the holes for the nostrils and the antor ...
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Jidapterus
''Jidapterus'' is a genus of chaoyangopterid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. The genus was in 2003 named by Dong Zhiming, Sun Yue-Wu and Wu Shao-Yuan. The type species is ''Jidapterus edentus''. The genus name is derived from ''Jílín Dàxué'' or "Jilin University" and a Latinized Greek ''pteron'', "wing". The specific name means "toothless" in Latin. Description ''Jidapterus'' is based on holotype CAD-01, a nearly complete skeleton with partial skull. The skull is toothless and relatively long, with a straight and very pointed beak, and a large hole where the antorbital fenestra is joined with the nostrils. The eye sockets are small, and there is no crest along the lower jaw as seen in ornithocheiroids, although a short projection was present at the back of the skull. The wingspan of this individual was estimated to be . Classification The classification of ''Jidapterus'' has been unstable; the original auth ...
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Apatorhamphus
''Apatorhamphus'' is an extinct genus of azhdarchoid pterosaur from the Kem Kem Group of Morocco. It might have been part of the Chaoyangopteridae. It is only known from a few snout fragments and it likely had a wingspan of between Discovery and naming During a visit in 2016 to the Tafilalt phosphate mine on the Aferdou N'Chaft plateau, near Hassi el Begaa, in Er Rachidia, British paleontologist David Michael Martill purchased the jaw fragment of a pterosaur from miners (specimen ''FSAC-KK 5010''). They had dug a tunnel in a thin fossil-containing layer on the edge of a quarry and finds were offered for sale there. These fossils belong to strata that belong to the Kem Kem Beds and date to the Cretaceous period, between the Albian to Cenomanian. In 2020, the new genus and species ''Apatorhamphus gyrostega'' was named and described by James McPhee, Nizar Ibrahim, Alex Kao, David M. Unwin, Roy Smith, and David M. Martill. McPhee ''et al.'' also referred various earlier finds to ...
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Eoazhdarcho
''Eoazhdarcho'' is a genus of azhdarchoid pterodactyloid pterosaur named in 2005 by Chinese paleontologists Lü Junchang and Ji Qiang. The type and only known species is ''Eoazhdarcho liaoxiensis''. The fossil was found in the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. Etymology The genus name, ''Eoazhdarcho'', combines a Greek ''eos'', "dawn" with the name of the genus ''Azhdarcho'', with the implication it was an early related form of the latter. The specific name, ''liaoxiensis'', refers to the ancient region Liaoxi. Description ''Eoazhdarcho'' is based on holotype GMN-03-11-02, a partial skeleton and lower jaw, and is distinguished from other pterosaurs by the proportions of its bones. The metacarpals are very elongated but the cervical vertebrae and hind limbs are not. It was relatively small by azhdarchoid standards, with a wingspan of about 1.6 meters (5.2 feet). Classification The describers first assigned ''Eoazhdarcho'' t ...
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Lacusovagus
''Lacusovagus'' (meaning "lake wanderer") is a genus of azhdarchoid pterodactyloid pterosaur from the Lower Cretaceous of Brazil. It is based on SMNK PAL 4325, a partial upper jaw comprising sections of the skull in front of the eyes. This specimen was found in rocks of the Early Cretaceous-age (probably Aptian stage, about 120 million years ago) Nova Olinda Member of the Crato Formation. The skull was long, and unusually wide. The section in front of the combined nasal-antorbital fenestra was relatively short. Also unusual was the combination of its toothless jaws and no bony head crest. ''Lacusovagus'' was described in 2008 by Mark Witton. The type species is ''L. magnificens'', meaning "grand lake wanderer", in reference to its large size—it is currently the largest pterosaur known from the Crato Formation with an estimated wingspan of approximately and a body mass of . ''Lacusovagus'' shares many characteristics with the basal azhdarchoid family Chaoyangopteridae, an ...
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Xericeps
''Xericeps'' is a genus of pterosaur from the Late Cretaceous Kem Kem Beds (which date to the late Albian or Cenomanian age) of southeastern Morocco. The name ''Xericeps'' comes from the grc, ξερός - meaning dry, referencing the Sahara Desert, in which the pterosaur was first found, and the la, cep from ''capere'', meaning "to catch" - alluding to the creature's forceps-like beak. Description ''Xericeps'' is a medium-sized edentulous (toothless) pterosaur. The term 'medium-sized', in the context of pterosaurs, is generally used to describe pterosaurs with a wingspan of 3–8 metres, and it is likely that ''Xericeps'' was nearer the lower end of this range. The holotype specimen is a partial anterior lower jaw, likely broken off right anterior to where the mandibular rami diverged. The jaw is upturned, with the occluding surface curved in lateral view. On the dorsal surface of the mandibular symphysis are a pair of ridges, similar to those seen in ''Alanqa'' and ''Argentin ...
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Liaoning
Liaoning () is a coastal province in Northeast China that is the smallest, southernmost, and most populous province in the region. With its capital at Shenyang, it is located on the northern shore of the Yellow Sea, and is the northernmost coastal province of the People's Republic of China. Historically a gateway between China proper and Manchuria, the modern Liaoning province was established in 1907 as Fengtian or Fengtien province and was renamed Liaoning in 1929. It was also known at that time as Mukden Province for the Manchu name of ''Shengjing'', the former name of Shenyang. Under the Japanese-puppet Manchukuo regime, the province reverted to its 1907 name, but the name Liaoning was restored for a brief time in 1945 and then again in 1954. Liaoning borders the Yellow Sea ( Korea Bay) and Bohai Sea in the south, North Korea's North Pyongan and Chagang provinces in the southeast, Jilin to the northeast, Hebei to the southwest, and Inner Mongolia to the northwest. The ...
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Antorbital Fenestra
An antorbital fenestra (plural: fenestrae) is an opening in the skull that is in front of the eye sockets. This skull character is largely associated with archosauriforms, first appearing during the Triassic Period. Among extant archosaurs, birds still possess antorbital fenestrae, whereas crocodylians have lost them. The loss in crocodylians is believed to be related to the structural needs of their skulls for the bite force and feeding behaviours that they employ.Preushscoft, H., Witzel, U. 2002. Biomechanical Investigations on the Skulls of Reptiles and Mammals. Senckenbergiana Lethaea 82:207–222.Rayfield, E.J., Milner, A.C., Xuan, V.B., Young, P.G. 2007. Functional Morphology of Spinosaur "Crocodile Mimic" Dinosaurs. JVP. 27(4):892–901. In some archosaur species, the opening has closed but its location is still marked by a depression, or fossa, on the surface of the skull called the antorbital fossa. The antorbital fenestra houses a paranasal sinus that is confluent with ...
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Azhdarchoid Mandibles
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutionary ...
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