Xericeps
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''Xericeps'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Kem Kem Beds The Kem Kem Group (commonly known as the Kem Kem beds) is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations, ...
(which date to the late
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
or
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
age) of southeastern
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
. The name ''Xericeps'' comes from the grc, ξερός - meaning dry, referencing the
Sahara Desert , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
, in which the
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
was first found, and the la, cep from ''capere'', meaning "to catch" - alluding to the creature's forceps-like beak.


Description

''Xericeps'' is a medium-sized edentulous (toothless) pterosaur. The term 'medium-sized', in the context of pterosaurs, is generally used to describe pterosaurs with a wingspan of 3–8 metres, and it is likely that ''Xericeps'' was nearer the lower end of this range. The holotype specimen is a partial anterior lower jaw, likely broken off right anterior to where the mandibular rami diverged. The jaw is upturned, with the occluding surface curved in lateral view. On the dorsal surface of the mandibular symphysis are a pair of ridges, similar to those seen in ''
Alanqa ''Alanqa'' is a genus of pterodactyloid pterosaur from the Late Cretaceous period ( Cenomanian stage) of what is now the Kem Kem Beds of southeastern Morocco. The name ''Alanqa'' comes from the Arabic word العنقاء ''al- ‘Anqā’'', for ...
'' and ''
Argentinadraco ''Argentinadraco'' (meaning "Argentina dragon") is an extinct genus of azhdarchoid pterosaur from the Late Cretaceous Portezuelo Formation of Argentina. It contains a single species, ''A. barrealensis'', named in 2017 by Alexander Kellner and Jo ...
''.


Classification

A 2021 study focused on ''
Aerotitan ''Aerotitan'' is a genus of large azhdarchid pterosaur known from the Late Cretaceous period (Maastrichtian stage) of what is now the Allen Formation of the Neuquén Basin in northern Patagonia, Argentina. Discovery and naming The type species ...
'' recovers it as a chaoyangopterid closely related to ''
Argentinadraco ''Argentinadraco'' (meaning "Argentina dragon") is an extinct genus of azhdarchoid pterosaur from the Late Cretaceous Portezuelo Formation of Argentina. It contains a single species, ''A. barrealensis'', named in 2017 by Alexander Kellner and Jo ...
''.Pêgas, R.V.; Holgado, B.; Ortiz David, L.D.; Baiano, M.A.; Costa, F.R. (August 21, 2021). "On the pterosaur Aerotitan sudamericanus (Neuquén Basin, Upper Cretaceous of Argentina), with comments on azhdarchoid phylogeny and jaw anatomy". Cretaceous Research. in press: Article 104998. doi:10.1016/j.cretres.2021.104998.


Discovery and naming

The holotype specimen - FSAC-KK-10700 was discovered by local mine workers at Aferdou N'Chaft, a small mesa near the oasis village of Hassi el Begaa in the
Errachidia Province Errachidia (Berber language: Imetɣaren) is a province located in eastern central Morocco in the region of Dra-Tafilalt. Errachidia is bordered by Algeria to the southeast and by the Moroccan provinces of Figig to the northeast, Midelt to the no ...
in south eastern Morocco on the Algerian border, and consists only of the pterosaur's fragmented jaws. The specimen was purchased directly at the mine site by British palaeontologist David M. Martill in January 2017, and thus it is possible to confidently establish its precise locality and
stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostrati ...
horizon. It is believed that ''Xericeps'' lived in the mid-
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
around the
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
-
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
ages (93.9-113.0 Ma). The holotype specimen's species epithet - "curvirostris" - comes from the Latin ''curvus'', meaning "curved" and ''rostrum'' meaning snout, or muzzle. This is due to the specimen's noticeably upward-curved beak.


See also

*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ...
*
List of pterosaurs This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered inva ...
*
2018 in paleontology Flora Plants Fungi Cnidarians Research * New three dimensionally phosphatized microfossils of coronate scyphozoan '' Qinscyphus necopinus'', including a new type of fossil embryo, are described from the Cambrian (Fortunian) Kuanchuanpu F ...


References

{{Portal bar, Paleontology, Cretaceous, Morocco Early Cretaceous pterosaurs Chaoyangopterids Early Cretaceous reptiles of Africa Late Cretaceous pterosaurs of Africa Fossil taxa described in 2018 Pterosaurs of Africa Albian genus first appearances Cenomanian genus extinctions