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Cardueae
The Cardueae are a tribe of flowering plants in the daisy family (Asteraceae) and the subfamily Carduoideae. Most of them are commonly known as thistles; four of the best known genera are '' Carduus'', '' Cynara'' (containing the widely eaten artichoke), ''Cirsium'', and ''Onopordum''. They are annual, biennial, or perennial herbs. Many species are thorny on leaves, stems, or involucre, and some have laticifers or resin conduits. Almost 80 genera comprising 2500 species are assigned to this tribe, native of temperate regions of Europe and Asia (especially the Mediterranean region and Minor Asia), Australia and tropical Africa; only three genera contain species native to the Americas.Bremer 1994 ''Asteraceae'': Cladistic and Classification ribe ''Carduae'': 112-156/ref> Taxonomy Cardueae is a synonym for Cynareae, but the name Cynareae was published almost a decade earlier, so has precedence. Some authors have divided the plants traditionally held to be in this tribe into th ...
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Thistle
Thistle is the common name of a group of flowering plants characterised by leaves with sharp prickles on the margins, mostly in the family Asteraceae. Prickles can also occur all over the planton the stem and on the flat parts of the leaves. These prickles are an adaptation that protects the plant from being eaten by herbivores. Typically, an involucre with a clasping shape similar to a cup or urn subtends each of a thistle's flower heads. The comparative amount of spininess varies dramatically by species. For example, ''Cirsium heterophyllum'' has minimal spininess while ''Cirsium spinosissimum'' is the opposite. Typically, species adapted to dry environments have greater spininess. The term thistle is sometimes taken to mean precisely those plants in the tribe Cardueae (synonym: Cynareae), especially the genera '' Carduus'', ''Cirsium'', and ''Onopordum''. However, plants outside this tribe are sometimes called thistles. Biennial thistles are particularly noteworthy for ...
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Cousinia
The genus ''Cousinia'' of the tribe Cardueae is in its current circumscription one of the larger genera in the Asteraceae, with approximately 650-700 species distributed in central and western Asia. Many of the species in this genus were once classified in genus ''Arctium ''Arctium'' is a genus of biennial plants commonly known as burdock, family Asteraceae. Native to Europe and Asia, several species have been widely introduced worldwide. Burdock's clinging properties, in addition to providing an excellent mecha ...'' (the burdocks). Species References External links USDA GRIN Cynareae Asteraceae genera {{Cynareae-stub ...
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Onopordum
''Onopordum'', or cottonthistle, is a genus of plants in the tribe Cardueae within the family Asteraceae. They are native to southern Europe, northern Africa, the Canary Islands, the Caucasus, and southwest and central Asia. They grow on disturbed land, roadsides, arable land and pastures. They are biennials (rarely short-lived perennials) with branched, spinose winged stems, growing 0.5–3 m tall. In the first season they form a basal rosette of gray-green felted leaves and rarely a few flower heads. In the second season they grow rapidly to their final height, flowering extensively, and then die off after seed maturation. The leaves are dentate or shallowly lobed to compound with several pinnatifid or deeply cut leaflets, and strongly spiny. The terminal flower head is typical for thistles, a semi-spherical to ovoid capitulum with purple (seldom white or pink) disc florets. There are no ray florets. The receptacle is glabrous with dentate margins. The tube of the corolla i ...
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Berardia
''Berardia'' is a monotypic genus of flowering plants in the family Asteraceae, containing the single species ''Berardia subacaulis''. It is an alpine plant native to the Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ....Garcia-Jacas, N., et al. (2002)Tribal and subtribal delimitation and phylogeny of the Cardueae (Asteraceae): a combined nuclear and chloroplast DNA analysis.''Molecular Phylogenetics and Evolution'' 22(1), 51-64. References Cynareae Flora of the Alps Monotypic Asteraceae genera {{Cynareae-stub ...
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Berardiinae (plant)
''Berardia'' is a monotypic genus of flowering plants in the family Asteraceae, containing the single species ''Berardia subacaulis''. It is an alpine plant native to the Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ....Garcia-Jacas, N., et al. (2002)Tribal and subtribal delimitation and phylogeny of the Cardueae (Asteraceae): a combined nuclear and chloroplast DNA analysis.''Molecular Phylogenetics and Evolution'' 22(1), 51-64. References Cynareae Flora of the Alps Monotypic Asteraceae genera {{Cynareae-stub ...
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Cynara
''Cynara'' is a genus of thistle-like perennial plants in the family Asteraceae. They are native to the Mediterranean region, the Middle East, northwestern Africa, and the Canary Islands. The genus name comes from the Greek ''kynara'', which means "artichoke".''Cynara''.
Flora of North America.
Among the better known species in this genus include: * '''' is the cardoon, artichoke thistle, or wild artichoke. The stems of cultivated varieties are used as food around the Mediterranean. It is a common source of a coagulant used as an alternative to in the manufac ...
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Carduus
''Carduus'' is a genus of flowering plants in the family Asteraceae, and the tribe Cardueae, one of two genera considered to be true thistles, the other being ''Cirsium''. Plants of the genus are known commonly as plumeless thistles.''Carduus''.
Integrated Taxonomic Information System (ITIS).
''Carduus''.
Flora of North America.
They are native to Eurasia and Africa, and several are known elsewhere as . This genus is noted for its disproportionately high number of

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Herbaceous Plant
Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of the ''Shorter Oxford English Dictionary'' defines "herb" as: #"A plant whose stem does not become woody and persistent (as in a tree or shrub) but remains soft and succulent, and dies (completely or down to the root) after flowering"; #"A (freq. aromatic) plant used for flavouring or scent, in medicine, etc.". (See: Herb) The same dictionary defines "herbaceous" as: #"Of the nature of a herb; esp. not forming a woody stem but dying down to the root each year"; #"BOTANY Resembling a leaf in colour or texture. Opp. scarious". Botanical sources differ from each other on the definition of "herb". For instance, the Hunt Institute for Botanical Documentation includes the condition "when persisting over more than one growing season, the parts o ...
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Thorn (botany)
In plant morphology, thorns, spines, and prickles, and in general spinose structures (sometimes called ''spinose teeth'' or ''spinose apical processes''), are hard, rigid extensions or modifications of leaves, roots, stems or buds with sharp, stiff ends, and generally serve the same function: physically deterring animals from eating the plant material. Description In common language the terms are used more or less interchangeably, but in botanical terms, thorns are derived shoots (so that they may or may not be branched, they may or may not have leaves, and they may or may not arise from a bud),Simpson, M. G. 2010. "Plant Morphology". In: ''Plant Systematics, 2nd. edition''. Elsevier Academic Press. Chapter 9.Judd, Campbell, Kellogg, Stevens, Donoghue. 2007. "Structural and Biochemical Characters". In: ''Plant Systematics, a phylogenetic approach, third edition''. Chapter 4. spines are derived from leaves (either the entire leaf or some part of the leaf that has vascular bundles ...
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Resin
In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on naturally occurring resins. Plants secrete resins for their protective benefits in response to injury. The resin protects the plant from insects and pathogens. Resins confound a wide range of herbivores, insects, and pathogens, while the volatile phenolic compounds may attract benefactors such as parasitoids or predators of the herbivores that attack the plant. Composition Most plant resins are composed of terpenes. Specific components are alpha-pinene, beta-pinene, delta-3 carene, and sabinene, the monocyclic terpenes limonene and terpinolene, and smaller amounts of the tricyclic sesquiterpenes, longifolene, caryophyllene, and delta-cadinene. Some resins also contain a high proportion of resin acids. Rosins on the other hand are less ...
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Involucral Bract
In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of a different color, shape, or texture. Typically, they also look different from the parts of the flower, such as the petals or sepals. A plant having bracts is referred to as bracteate or bracteolate, while one that lacks them is referred to as ebracteate and ebracteolate, without bracts. Variants Some bracts are brightly-coloured and serve the function of attracting pollinators, either together with the perianth or instead of it. Examples of this type of bract include those of ''Euphorbia pulcherrima'' (poinsettia) and ''Bougainvillea'': both of these have large colourful bracts surrounding much smaller, less colourful flowers. In grasses, each floret (flower) is enclosed in a pair of papery bracts, called the lemma (lower bract) and pa ...
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Latex
Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosperms). It is a complex emulsion that coagulates on exposure to air, consisting of proteins, alkaloids, starches, sugars, oils, tannins, resins, and gums. It is usually exuded after tissue injury. In most plants, latex is white, but some have yellow, orange, or scarlet latex. Since the 17th century, latex has been used as a term for the fluid substance in plants, deriving from the Latin word for "liquid". It serves mainly as defense against herbivorous insects. Latex is not to be confused with plant sap; it is a distinct substance, separately produced, and with different functions. The word latex is also used to refer to natural latex rubber, particularly non-vulcanized rubber. Such is the case in products like latex gloves, latex condoms ...
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