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Carabidae
Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal families. They belong to the Adephaga. Members of the family are primarily carnivorous, but some members are phytophagous or omnivorous. Description and ecology Although their body shapes and coloring vary somewhat, most are shiny black or metallic and have ridged wing covers (elytra). The elytra are fused in some species, particularly the large Carabinae, rendering the beetles unable to fly. The species ''Mormolyce phyllodes'' is known as violin beetle due to their peculiarly shaped elytra. All carabids except the quite primitive flanged bombardier beetles (Paussinae) have a groove on their fore leg tibiae bearing a comb of hairs used for cleaning their antennae. Defensive secretions Typical for the ancient beetle suborder Adephaga to ...
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Harpalinae
Harpalinae is a huge subfamily of ground beetles that contains 20,000 species or ~6,400 spp. in 24 tribes worldwide, according to others. A rarely used common name for the subfamily is the harp beetles. The Harpalinae contain the most apomorphic ground beetles, displaying a wide range of forms and behaviors. Some are, rare among ground beetles, omnivores or even herbivores. Many closely related subfamilies have been treated as subordinate taxa of the Harpalinae by various authors. Among these are the Dryptinae, Lebiinae (including Cyclosominae, Mormolycinae, Odacanthinae, Perigoninae), Licininae (including Chlaeniinae, Oodinae), Orthogoniinae, Panagaeinae, Platyninae, Pseudomorphinae, Pterostichinae (including Zabrinae). Here, they are considered independent families within the harpaline (''sensu lato'') assemblage, and this is also tentatively assumed for the enigmatic monotypic genus ''Ginema''. Systematics At least 4 large and several smaller supertribes can be dist ...
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Cicindelinae
Tiger beetles are a family of beetles, Cicindelidae, known for their aggressive predatory habits and running speed. The fastest known species of tiger beetle, ''Rivacindela hudsoni'', can run at a speed of , or about 125 body lengths per second. As of 2005, about 2,600 species and subspecies were known, with the richest diversity in the Oriental (Indo-Malayan) region, followed by the Neotropics. While historically treated as a subfamily of ground beetles (Carabidae) under the name Cicindelinae, several studies since 2020 indicated that they should be treated as a family, the Cicindelidae, which are a sister group to Carabidae within the Adephaga. Description Tiger beetles often have large bulging eyes, long, slender legs and large curved mandibles. All are predatory, both as adults and as larvae. The genus ''Cicindela'' has a cosmopolitan distribution. Other well-known genera include ''Tetracha'', ''Omus'', ''Amblycheila'' and '' Manticora''. While members of the genus ''Cicindela ...
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Nototylinae
Nototylinae is a subfamily of beetles in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal fami .... It contains the single genus ''Nototylus'' with two species, '' Nototylus fryi'' and '' Nototylus balli'' which was described in 2020. The first described species ''fryi'' was represented by a single specimen from Espiritu Santo where the forests were converted for plantations of sugarcane or cacao, and for cattle rearing. The species is considered likely extinct. References Monotypic insect taxa Carabidae subfamilies {{Carabidae-stub ...
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Pseudomorphinae
Pseudomorphinae is a subfamily of ground beetles (family Carabidae). Occasionally it was treated as a tribe Pseudomorphini of subfamily Harpalinae, particularly when this was circumscribed loosely. It contains the following genera: * ''Adelotopus'' Hope, 1834 * ''Cainogenion'' Notman, 1925 * ''Cryptocephalomorpha'' Ritsema, 1875 * '' Guyanemorpha'' Terry L. Erwin, 2013 * ''Paussotropus'' Waterhouse, 1877 * ''Protoscalidion'' Schaufuss, 1888 * ''Pseudomorpha'' Kirby, 1825 * ''Sphallomorpha ''Sphallomorpha'' is a genus of beetles in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in E ...'' Westwood, 1837 References Carabidae subfamilies {{Carabidae-stub ...
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Gineminae
''Ginema'' is a genus of ground beetles (family Carabidae). The single described species, ''Ginema thomasi'', is known only from Bolivia. Its closest living relatives are unknown, and is sometimes separated as subfamily Gineminae. While this species certainly does not resemble any other tribe of ground beetles, it is not yet fully resolved whether treatment as a monotypic subfamily adequately expresses its phylogenetic position. Alternatively, Gineminae can be downranked to a tribe Ginemini within subfamily Harpalinae Harpalinae is a huge subfamily of ground beetles that contains 20,000 species or ~6,400 spp. in 24 tribes worldwide, according to others. A rarely used common name for the subfamily is the harp beetles. The Harpalinae contain the most apomorphic ... (particularly when these are loosely circumscribed) or a related lineage. References Monotypic Carabidae genera Carabidae {{Carabidae-stub ...
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Hiletinae
Hiletinae is a subfamily of beetles in the family Carabidae, containing 21 species in two genera. All of the species in the genus '' Hiletus'', as well as 6 species in '' Eucamaragnathus'', live in Africa. The other species in ''Eucamaragnathus'' live in either Southeast Asia and India, or in South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout .... Classification * Genus '' Eucamaragnathus'' Jeannel, 1938 ** '' Eucamaragnathus amapa'' Erwin & Stork, 1985 ** '' Eucamaragnathus angulicollis'' (Jeannel, 1938) ** '' Eucamaragnathus aterrimus'' (Jeannel, 1938) ** '' Eucamaragnathus batesii'' (Chaudoir, 1861) ** '' Eucamaragnathus bocandei'' (Alluaud, 1914) ** '' Eucamaragnathus borneensis'' Erwin & Stork, 1985 ** '' Eucamaragnathus brasiliensis'' (Negre, 1966) ** '' Eucamarag ...
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Elaphrinae
Elaphrinae is a subfamily of beetles in the family Carabidae, containing the following genera: * '' Blethisa'' Bonelli, 1810 * ''Diacheila'' Motschulsky, 1844 * ''Elaphrus ''Elaphrus'' is a genus of ground beetle native to the Palearctic, the Nearctic, the Near East and Northern Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singula ...'' Fabricius, 1775 References Carabidae subfamilies {{Carabidae-stub ...
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Pterostichinae
Pterostichinae is a subfamily of ground beetles (family Carabidae). It belongs to the advanced harpaline assemblage, and if these are circumscribed ''sensu lato'' as a single subfamily, Pterostichinae are downranked to a tribe Pterostichini. However, as the former Pterostichitae supertribe of the Harpalinae as loosely circumscribed does seem to constitute a lineage rather distinct from '' Harpalus'', its core group is here considered to be the present subfamily and the Harpalinae are defined more narrowly. They are usually mid-sized and rather stout ground beetles. Coloration is typically dark and without conspicuous patterns, but often with a strong sheen like polished metal. They are widely distributed and inhabit a wide range of terrestrial habitats. Unlike the more basal ground beetles which only eat small animals, the Pterostichinae include a large proportion of omnivorous or even herbivorous taxa. Systematics This group includes the following tribes and genera: Tribe A ...
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Loricerinae
Loricerinae is a subfamily of beetles in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal fami .... It contains the single genus ''Loricera'' with the following species: * '' Loricera aptera'' Ball & Erwin, 1969 * '' Loricera balli'' Sciaky & Facchini, 1999 * '' Loricera barbarae'' Sciaky & Facchini, 1999 * '' Loricera decempunctata'' Eschscholtz, 1833 * '' Loricera foveata'' Leconte, 1851 * '' Loricera kryzhanowskiji'' Sciaky & Facchini, 1999 * '' Loricera mirabilis'' Jedlicka, 1932 * '' Loricera obsoleta'' Semenov, 1889 * '' Loricera ovipennis'' Semenov, 1889 * '' Loricera pilicornis'' Fabricius, 1775 * '' Loricera rotundicollis'' Chaudoir, 1863 * '' Loricera stevensi'' Andrewes, 1920 * '' Loricera wollastonii'' Javet, 1852 References Carabidae subfamili ...
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Promecognathinae
Promecognathinae is a subfamily of beetles in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal fami .... It contains 10 species in 7 genera: * Tribe Dalyatini Mateu, 2002 ** Genus '' Dalyat'' Mateu, 2002 *** '' Dalyat mirabilis'' Mateu, 2002 * Tribe Palaeoaxinidiini McKay, 1991 ** Genus '' Palaeoaxinidium'' McKay, 1991 *** '' Palaeoaxinidium orapense'' McKay, 1991 * Tribe Promecognathini LeConte, 1853 ** Genus '' Axinidium'' Sturm, 1843 *** '' Axinidium africanum'' Sturm, 1843 *** '' Axinidium angulatum'' Basilewsky, 1963 ** Genus '' Holaxinidium'' Basilewsky, 1963 *** '' Holaxinidium fitsimonsi'' Basilewsky, 1963 ** Genus '' Metaxinidium'' Basilewsky, 1963 *** '' Metaxinidium leleupi'' Basilewsky, 1963 *** '' Metaxinidium nanum'' Basilewsky, 1963 ** Genus '' Parax ...
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Protorabinae
Protorabinae is a subfamily of beetles in the family Carabidae. It contains 31 species in 10 genera: * Genus '' Aethocarabus'' Ren, 1995 ** '' Athocarabus levigata'' Ren, 1995 * Genus '' Cordorabus'' Ponomarenko, 1977 ** '' Cordorabus antennatus'' Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/ Oxfordian ** '' Cordorabus minimus'' Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian ** '' Cordorabus notatus'' Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian ** '' Cordorabus striatus'' Ponomarenko, 1986 Gurvan-Eren Formation, Mongolia, Aptian ** '' Cordorabus vittatus'' Ponomarenko, 1980 Mogotuin Formation, Mongolia, Aptian * Genus '' Cretorabus'' Ponomarenko, 1977 ** '' Cretorabus capitatus'' Ponomarenko, 1977 Zaza Formation, Russia, Aptian ** '' Cretorabus latus'' Ponomarenko, 1977 Zaza Formation, Russia, Aptian ** '' Cretorabus orientalis'' Ponomarenko, 1989 Tsagaantsav Formation, Mongolia, Valanginian ** '' Cretorabus ...
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Apotominae
Apotominae is a subfamily of beetles in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal fami .... It contains the single genus ''Apotomus'' with the following species: * '' Apotomus alluaudi'' Jeannel, 1946 * '' Apotomus angusticollis'' J. Muller, 1943 * '' Apotomus annulaticornis'' Peringuey, 1896 * '' Apotomus atripennis'' Motschulsky, 1858 * '' Apotomus australis'' Castelnau, 1867 * '' Apotomus chaudoirii'' Wollaston, 1860 * '' Apotomus clypeonitens'' J. Muller, 1943 * '' Apotomus fairmairei'' Jeannel, 1946 * '' Apotomus flavescens'' Apetz, 1854 * '' Apotomus hirsutulus'' Bates, 1892 * '' Apotomus latigena'' Reitter, 1892 * '' Apotomus minor'' Baehr, 1990 * '' Apotomus neghellianus'' G. Muller, 1942 * '' Apotomus qiongshanensis'' Tian, 2000 * '' Apotomus reichardti ...
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