Protorabinae is a subfamily of
beetle
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s in the family
Carabidae
Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal fami ...
. It contains 31 species in 10 genera:
* Genus ''
Aethocarabus''
Ren, 1995
** ''
Athocarabus levigata''
Ren, 1995
* Genus ''
Cordorabus''
Ponomarenko, 1977
** ''
Cordorabus antennatus''
Ponomarenko, 1977 Karabastau Formation
The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that ...
, Kazakhstan, Callovian
In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
/ Oxfordian
** ''
Cordorabus minimus''
Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** ''
Cordorabus notatus''
Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** ''
Cordorabus striatus''
Ponomarenko, 1986 Gurvan-Eren Formation, Mongolia, Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
** ''
Cordorabus vittatus''
Ponomarenko, 1980 Mogotuin Formation, Mongolia, Aptian
* Genus ''
Cretorabus''
Ponomarenko, 1977
** ''
Cretorabus capitatus''
Ponomarenko, 1977 Zaza Formation
The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous period. It is Aptian in age and consists of Sandstone, sandstones, Siltstone, siltstones, Marl, marls and bituminous Shale, shales, deposi ...
, Russia, Aptian
** ''
Cretorabus latus''
Ponomarenko, 1977 Zaza Formation, Russia, Aptian
** ''
Cretorabus orientalis''
Ponomarenko, 1989 Tsagaantsav Formation
The Tsagaantsav Formation, Tsagantsab Formation or Tsagan-Tsab Formation (Russian: Tsagaantsav Svita) is an Early Cretaceous (Hauterivian to Barremian) geologic formation in Mongolia.[Valanginian
In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretaceou ...]
** ''
Cretorabus ovalis''
Ponomarenko, 1989 Dzun-Bain Formation, Mongolia, Aptian
** ''
Cretorabus rasnitsyni''
Wang & Zhang, 2011 Yixian Formation
The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ex ...
, China, Aptian
** ''
Cretorabus sulcatus''
Ponomarenko, Coram & Jarzembowski, 2005 Lulworth Formation
The Lulworth Formation is a Formation (geology), geologic formation in England. It dates from the late Tithonian to the mid Berriasian. It is a subunit of the Purbeck Group. In Dorset, it consists of three members, which are in ascending order, t ...
, United Kingsom, Berriasian
In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ...
* Genus ''
Lithorabus''
Ponomarenko, 1977
** ''
Lithorabus incertus''
Ponomarenko, 1977 Dzhil Formation, Kyrgyzstan Hettangian
The Hettangian is the earliest age and lowest stage of the Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (My ...
* Genus ''
Nebrorabus''
Ponomarenko, 1989
** ''
Nebrorabus baculum''
Ponomarenko, 1989
** ''
Nebrorabus capitatus''
Ponomarenko, 1989
** ''
Nebrorabus elongatus''
Ponomarenko, 1986
** ''
Nebrorabus nebrioides''
Ponomarenko, 1989
** ''
Nebrorabus tumoculus''
Ren, 1995
* Genus ''
Obesofemoria''
Hong, 1982
** ''
Obesofemoria chijingiaoensis''
Hong, 1982
* Genus ''
Ovrabites''
Ponomarenko, 1977
** ''
Ovrabites incertus''
Ponomarenko, 1993 Emanra Formation, Russia, Turonian
The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by t ...
** ''
Ovrabites jurassicus''
Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** ''
Ovrabites ovalis''
Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
* Genus ''
Penecupes''
Ren, 1995
** ''
Penecupes rapax''
Ren, 1995 Lushangfen Formation, China, Aptian
* Genus ''
Protorabus''
Ponomarenko, 1977
** ''
Protorabus crassus''
Ponomarenko, 1989 Daya Formation, Russia, Hauterivian
The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...
** ''
Protorabus kobdoensis''
Ponomarenko, 1986 Gurvan-Eren Formation, Mongolia, Aptian
** ''
Protorabus magnus''
Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** ''
Protorabus nigrimonticola''
Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** ''
Protorabus planus''
Ponomarenko, 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
** ''
Protorabus polyphlebius''
Ren, 1995 Lushangfen Formation, China, Aptian
** ''
Protorabus tsaganensis''
Ponomarenko, 1989 Dzun-Bain Formation, Mongolia, Aptian
* Genus ''
Sinocarabus''
Hong, 1982
** ''
Sinocarabus leptoceroides''
Hong, 1982
References
Carabidae subfamilies
{{Carabidae-stub