Brachaucheninae
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Brachaucheninae
Thalassophonea is an extinct clade of pliosaurids from the Middle Jurassic to the early Late Cretaceous (Callovian to Turonian) of Australia, Europe, North America and South America. ''Thalassophonea'' was erected by Roger Benson and Patrick Druckenmiller in 2013. The name is derived from Greek ''thalassa'' (θάλασσα), "sea", and ''phoneus'' (φονεύς), "murderer". It is a stem-based taxon defined as "all taxa more closely related to ''Pliosaurus brachydeirus'' than to ''Marmornectes candrewi''". It includes the short necked and large headed taxa that typify the family. Classification The following cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... follows an analysis by Benson & Druckenmiller (2014). References Middle Jurassic first appearances Cretac ...
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Megacephalosaurus
''Megacephalosaurus'' (; "great-headed lizard") is an extinct genus of short-necked pliosaur that inhabited the Western Interior Seaway of North America about 94 to 93 million years ago during the Turonian stage of the Late Cretaceous, containing the single species ''M. eulerti''. It is named after its large head, which is the largest of any plesiosaur in the continent and measures up to in length. ''Megacephalosaurus'' was one of the largest marine reptiles of its time with an estimated length of . Its long snout and consistently sized teeth suggest that it preferred a diet on smaller-sized prey. Remains representing the pliosaur include two fossil skulls, three ribs, and a neural arch. The fossils have been found in deposits of the Carlile Shale and Greenhorn Limestone in Kansas and elsewhere in the midwestern United States. First discovered in 1950, they were originally thought to have been giant fossils of a closely related pliosaur known as '' Brachauchenius lucasi''. But b ...
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Luskhan
''Luskhan'' (meaning "water spirit chief") is an extinct genus of brachauchenine pliosaur from the Cretaceous of Russia. The type and only species is ''Luskhan itilensis'', named by Valentin Fischer and colleagues in 2017 from a well-preserved and nearly complete skeleton. As an early-diverging brachauchenine, ''Luskhan'' consequently exhibits an intermediate combination of traits seen in more basal (less specialized) and more derived (more specialized) pliosaurs. However, ''Luskhan'' departs significantly from other pliosaurs in that it exhibits a lack of adaptations in its skull to feeding on large prey; its slender snout, small teeth, and short tooth rows instead indicate a skull adapted for feeding on small, soft prey. With these features, it is the pliosaur that approaches closest to the distantly-related piscivorous polycotylids, having convergently evolved these traits more than 10 million years apart. Discovery and naming A nearly complete fossil skeleton of a pliosau ...
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Pliosaurid
Pliosauridae is a family of plesiosaurian marine reptiles from the Latest Triassic to the early Late Cretaceous ( Rhaetian to Turonian stages) of Australia, Europe, North America and South America. The family is more inclusive than the archetypal short-necked large headed species that are placed in the subclade Thalassophonea, with basal forms resembling other plesiosaurs with long necks. They became extinct during the early Late Cretaceous and were subsequently replaced by the mosasaurs. It was formally named by Harry G. Seeley in 1874. Relationships Pliosauridae is a stem-based taxon defined in 2010 (and in earlier studies in a similar manner) as "all taxa more closely related to ''Pliosaurus brachydeirus'' than to '' Leptocleidus superstes'', '' Polycotylus latipinnis'' or ''Meyerasaurus victor''". The family Brachauchenidae has been proposed to include pliosauroids which have very short necks and may include ''Brachauchenius'' and ''Kronosaurus''. However, modern cladisti ...
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Polyptychodon
''Polyptychodon'' (meaning 'shaped fin tooth') is a genus of pliosaurid found in Middle-Late Cretaceous marine deposits in southern England, France and Argentina. It has been considered a ''nomen dubium'' in a 2016 review. History of discovery The type species, ''P. interruptus'' is known from an isolated tooth from the Late Cretaceous Chalk Group of southern England. Owen described a second nominal species of the genus, ''P. continuus'', from an isolated tooth collected in the Hythe Formation of Maidstone, Kent. (The macronarian sauropod '' Dinodocus'' was mistakenly thought to be conspecific with ''P. continuus'' before it was correctly recognized as a dinosaur and not a plesiosaur.) Numerous pliosaurid teeth and vertebrae from England and eastern France have been previously assigned to ''Polyptychodon'', including isolated vertebrae from France which were misidentified as a sauropod. Comparison between Albian-age isolated vertebrae from marine deposits in France and ''Krono ...
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Pliosaurus
''Pliosaurus'' (meaning 'more lizard') is an extinct genus of thalassophonean pliosaurid known from the Kimmeridgian and Tithonian stages (Late Jurassic) of Europe and South America. Their diet would have included fish, cephalopods, and marine reptiles. This genus has contained many species in the past but recent reviews found only six (''P. brachydeirus'', ''P. carpenteri'', ''P. funkei'', ''P. kevani'', ''P. rossicus'' and ''P. westburyensis'') to be valid, while the validity of two additional species awaits a petition to the International Code of Zoological Nomenclature. Currently, ''P. brachyspondylus'' and ''P. macromerus'' are considered dubious, while ''P. portentificus'' is considered undiagnostic. Most species of ''Pliosaurus'' reached in length and in body mass, while ''P. rossicus'' and ''P. funkei'' may have reached or even exceeded in length and in body mass, being the largest plesiosaurs of all time. Species of this genus are differentiated from other pliosaurids ...
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Peloneustes
''Peloneustes'' (meaning "mud swimmer") is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of ''Plesiosaurus'' by palaeontologist Harry Govier Seeley in 1896, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to ''Peloneustes'', ''P. philarchus'' is currently the only one still considered valid, with the others moved to different genera, considered '' nomina dubia'', or synonymised with ''P. philarchus''. Some of the material formerly assigned to ''P. evansi'' have since been reassigned to ''"Pliosaurus" andrewsi''. ''Peloneustes'' is known from many specimens, including some very complete material. With a total length of , ''Peloneustes'' is not a large pliosaurid. It had a large, triangular skull, which occupied about a fifth of its body length. The ...
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Liopleurodon
''Liopleurodon'' (; meaning 'smooth-sided teeth') is an extinct genus of large, carnivorous marine reptile belonging to the Thalassophonea, a clade of short-necked pliosaurid plesiosaurs. ''Liopleurodon'' lived from the Callovian Stage of the Middle Jurassic to the Kimmeridgian stage of the Late Jurassic Period (c. 166 to 155 mya). It was the apex predator of the Middle to Late Jurassic seas that covered Europe. The largest species, ''L. ferox'', is estimated to have grown up to in length, but could have been larger. The name "Liopleurodon" (meaning "smooth-sided tooth") derives from Ancient Greek words: ', "smooth"; ', "side" or "rib"; and ', "tooth". Discovery and species Even before ''Liopleurodon'' was named, material likely belonging to it was described. In 1841, Hermann von Meyer named the species ''Thaumatosaurus oolithicus'' based on a fragmentary specimen consisting of partial teeth, skull elements, vertebrae, and ribs from deposits in Württemberg, Germany poss ...
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Liopleurodon Ferox
''Liopleurodon'' (; meaning 'smooth-sided teeth') is an extinct genus of large, carnivorous marine reptile belonging to the Thalassophonea, a clade of short-necked pliosaurid plesiosaurs. ''Liopleurodon'' lived from the Callovian Stage of the Middle Jurassic to the Kimmeridgian stage of the Late Jurassic Period (c. 166 to 155 mya). It was the apex predator of the Middle to Late Jurassic seas that covered Europe. The largest species, ''L. ferox'', is estimated to have grown up to in length, but could have been larger. The name "Liopleurodon" (meaning "smooth-sided tooth") derives from Ancient Greek words: ', "smooth"; ', "side" or "rib"; and ', "tooth". Discovery and species Even before ''Liopleurodon'' was named, material likely belonging to it was described. In 1841, Hermann von Meyer named the species ''Thaumatosaurus oolithicus'' based on a fragmentary specimen consisting of partial teeth, skull elements, vertebrae, and ribs from deposits in Württemberg, Germany poss ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, smaller islands. With an area of , Australia is the largest country by area in Oceania and the world's List of countries and dependencies by area, sixth-largest country. Australia is the oldest, flattest, and driest inhabited continent, with the least fertile soils. It is a Megadiverse countries, megadiverse country, and its size gives it a wide variety of landscapes and climates, with Deserts of Australia, deserts in the centre, tropical Forests of Australia, rainforests in the north-east, and List of mountains in Australia, mountain ranges in the south-east. The ancestors of Aboriginal Australians began arriving from south east Asia approximately Early human migrations#Nearby Oceania, 65,000 years ago, during the Last Glacial Period, last i ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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