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Burkesuchus
''Burkesuchus'' (meaning "Coleman Burke's crocodile") is an extinct genus of basal neosuchians from the Upper Jurassic (Tithonian)-aged Toqui Formation of southern Chile. The genus is currently represented by a single species ''B. mallingrandensis'', named and described in 2021 on the basis of the holotype and additional referred specimens. ''Burkesuchus mallingrandensis'' was noticeably small being around an estimated 70 cm (27.5 inches) in length. The cranium is dorsoventrally depressed and transversely wide posteriorly and distinguished by a posterroventrally flexed wing-like squamosal. ''Burkesuchus'' was a small carnivore likely on invertebrates animals such as insects, crustaceans and may had feed on small aquatic vertebrates like fish. ''B''. ''mallingrandensis'' did not have the ability to feed on large prey items or tear large chunks of meat like modern day crocodiles do. The holotype SQO.PV 17700, consist of a cervical neural arch, partial neurocranium, four dorsal v ...
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Toqui Formation
The Toqui Formation is a geological formation in the Aysén Region of southern Chile. It has been dated to the Tithonian stage of the Late Jurassic by uranium–lead dating of zircons, providing an age of 147 ± 0.1 Ma. It consists of an sequence of clastic sedimentary sandstones and conglomerates, interbedded with volcanic tuffs and ignimbrite. The dinosaurs ''Chilesaurus'' and indeterminate diplodocids and the mesoeucrocodylian ''Burkesuchus'' are known from the formation. The formation was deposited in a fluvio-deltaic environment.Toqui Formation, Aysén
at Fossilworks.org


Paleobiota of the Toqui Formation


Crocodylomorphs


Dinosaurs


See al ...
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Neosuchia
Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all crocodylomorphs more closely related to ''Crocodylus niloticus'' (the Nile Crocodile) than to ''Notosuchus terrestris''. Members of Neosuchia generally share a crocodilian-like bodyform adapted to freshwater aquatic life, as opposed to the terrestrial habits of more basal crocodylomorph groups. The earliest neosuchian is suggested to be the Early Jurassic ''Calsoyasuchus'', which lived during the Sinemurian and Pliensbachian stages in North America. It is often identified as a member of Goniopholididae, though this is disputed, and the taxon may lie outside Neosuchia, which places the earliest records of the group in the Middle Jurassic. Characteristics A tooth notch between the maxilla and premaxilla is a basal characteristic of the Neosuchia, although it is lost in some more derived forms, most nota ...
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Neosuchia
Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all crocodylomorphs more closely related to ''Crocodylus niloticus'' (the Nile Crocodile) than to ''Notosuchus terrestris''. Members of Neosuchia generally share a crocodilian-like bodyform adapted to freshwater aquatic life, as opposed to the terrestrial habits of more basal crocodylomorph groups. The earliest neosuchian is suggested to be the Early Jurassic ''Calsoyasuchus'', which lived during the Sinemurian and Pliensbachian stages in North America. It is often identified as a member of Goniopholididae, though this is disputed, and the taxon may lie outside Neosuchia, which places the earliest records of the group in the Middle Jurassic. Characteristics A tooth notch between the maxilla and premaxilla is a basal characteristic of the Neosuchia, although it is lost in some more derived forms, most nota ...
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Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name "Malm" indicates rocks of Late Jurassic age. In the past, ''Malm'' was also used to indicate the unit of geological time, but this usage is now discouraged to make a clear distinction between lithostratigraphic and geochronologic/chronostratigraphic units. Subdivisions The Late Jurassic is divided into three ages, which correspond with the three (faunal) stages of Upper Jurassic rock: Paleogeography During the Late Jurassic Epoch, Pangaea broke up into two supercontinents, Laurasia to the north, and Gondwana to the south. The result of this break-up was the spawning of the Atlantic Ocean. However, at this time, the Atlantic Ocean was relatively narrow. Life forms of the epoch This epoch is well known for many famous types of dinosau ...
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Ulna
The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of the forearm as the little finger. It runs parallel to the radius, the other long bone in the forearm. The ulna is usually slightly longer than the radius, but the radius is thicker. Therefore, the radius is considered to be the larger of the two. Structure The ulna is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. It is broader close to the elbow, and narrows as it approaches the wrist. Close to the elbow, the ulna has a bony process, the olecranon process, a hook-like structure that fits into the olecranon fossa of the humerus. This prevents hyperextension and forms a hinge joint with the trochlea of the humerus. There is ...
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Alligatorium
''Alligatorium'' is an extinct genus of atoposaurid crocodylomorph from Late Jurassic marine deposits in France. Systematics The type species is ''A. meyeri'', named in 1871 from a single specimen from Cerin, eastern France. Two more nominal species, ''A. franconicum'', named in 1906, and ''A paintenense'', named in 1961, are based on now-missing specimens from Bavaria, southern Germany, and were synonymized into a single species, for which ''A. franconicum'' has priority. A 2016 review of Atoposauridae removed ''A. franconicum'' from ''Alligatorium'' and placed at Neosuchia ''incertae sedis''. ''Alligatorium depereti'', described in 1915, was reassigned to its own genus, ''Montsecosuchus ''Montsecosuchus'' is an extinct genus of atoposaurid crocodylomorphs. It is the replacement generic name for ''Alligatorium depereti'', which was described in 1915 from the Montsec Lithographic Limestone quarry of Spain. Fossils found from this ...'', in 1988. References Late Jura ...
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Peirosauridae
Peirosauridae is a Gondwanan family of mesoeucrocodylians that lived during the Cretaceous period. It was a clade of terrestrial crocodyliforms that evolved a rather dog-like form, and were terrestrial carnivores. It was phylogenetically defined in 2004 as the most recent common ancestor of ''Peirosaurus'' and Lomasuchinae and all of its descendants. Lomasuchinae is a subfamily of peirosaurids that includes the genus ''Lomasuchus''. Lomasuchinae was defined in the same 2004 study as the most recent common ancestor of ''Lomasuchus'' and Mahajangasuchini and all of its descendants. Mahajangasuchini, also constructed in the study, was defined as the most recent common ancestor of ''Mahajangasuchus'' and ''Uberabasuchus'' and all of its descendants. However, all more recent phylogenetic analyses placed ''Mahajangasuchus'' within its own family, Mahajangasuchidae, along with the newly named ''Kaprosuchus''. Genera The following list of genera follows Martinelli ''et al.'', 2012 unless ...
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Mahajangasuchidae
Mahajangasuchidae is an extinct family of notosuchian crocodyliforms. It currently contains two genera, '' Mahajangasuchus'' and '' Kaprosuchus'', both of which lived during the Late Cretaceous in Gondwana. It is defined as the most inclusive clade containing ''Mahajangasuchus insignis'' but not ''Notosuchus terrestris'', '' Simosuchus clarki'', '' Araripesuchus gomesii'', ''Baurusuchus pachecoi'', '' Peirosaurus torminni'', '' Goniopholis crassidens'', '' Pholidosaurus schaumbergensis'', or ''Crocodylus niloticus''. Phylogenetically, Mahajangasuchidae is placed just outside pholidosaurids and more derived neosuchians. Defining characters of the family include fused nasals, a jaw articulation below the posterior maxillary tooth row, a deep mandibular symphysis that is oriented anterodorsally, and the formation of a hornlike posterodorsal process from the squamosal and parietal (which is much more pronounced in ''Kaprosuchus''). Phylogeny Cladogram showing the phylogenetic ...
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Uruguaysuchidae
Uruguaysuchidae is a family of notosuchian crocodyliforms that lived in South America and Africa during the Cretaceous period. It includes the genera ''Araripesuchus ''Araripesuchus'' is a genus of extinct crocodyliform that existed during the Cretaceous period of the late Mesozoic era some 125 to 66 million years ago. Six species of ''Araripesuchus'' are currently known. They are generally considered to be n ...'' and '' Uruguaysuchus''. Below is a cladogram from Soto ''et al.'' (2011): References Early Cretaceous crocodylomorphs Terrestrial crocodylomorphs Late Cretaceous crocodylomorphs Early Cretaceous first appearances Late Cretaceous extinctions Prehistoric reptile families {{paleo-archosaur-stub ...
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Notosuchia
Notosuchia is a suborder of primarily Gondwanan mesoeucrocodylian crocodylomorphs that lived during the Jurassic and Cretaceous. Some phylogenies recover Sebecosuchia as a clade within Notosuchia, others as a sister group (see below); if Sebecosuchia is included within Notosuchia its existence is pushed into the Middle Miocene, about 11 million years ago. Fossils have been found from South America, Africa, Asia, and Europe. Notosuchia was a clade of terrestrial crocodilians that evolved a range of feeding behaviours, including herbivory ('' Chimaerasuchus''), omnivory (''Simosuchus''), and terrestrial hypercarnivory (''Baurusuchus''). It included many members with highly derived traits unusual for crocodylomorphs, including mammal-like teeth, flexible bands of shield-like body armor similar to those of armadillos (''Armadillosuchus''), and possibly fleshy cheeks and pig-like snouts (''Notosuchus''). The suborder was first named in 1971 by Zulma Gasparini and has since undergone ...
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Stolokrosuchus
''Stolokrosuchus'' is an extinct genus of neosuchian crocodylomorph that lived during the Early Cretaceous. Its fossils, including a skull with a long thin snout and bony knobs on the prefrontal, have been found in Niger. ''Stolokrosuchus'' was described in 2000 by Hans Larsson and Boubacar Gado. The type species is ''S. lapparenti''. They initially described it as related to Peirosauridae, if not a member of that family. One study has shown it to be related to ''Elosuchus''. However, more recent works usually find ''Stolokrosuchus'' to be one of the basalmost neosuchian, only distantly related to the elosuchid or pholidosaurid, ''Elosuchus ''Elosuchus'' is an extinct genus of neosuchian crocodyliform that lived during the Middle Cretaceous of what is now Africa (Niger, Morocco and Algeria). Description and taxonomy ''Elosuchus'' had an elongated snout like a gharial and was prob ...''. Phylogeny The cladogram following by Nicholl ''et al.'' 2021: References ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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