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Brachyopterella
''Brachyopterella'' is a genus of prehistoric eurypterid classified as part of the Rhenopteridae family. Two species, both from the Silurian period, are known; ''B. pentagonalis'' from Norway and ''B. ritchei'' from Scotland.Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 16.0 http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf (PDF). Description ''Brachyopterella'' was a small stylonurine eurypterid with a pentagonal carapace with little to no marginal rim and a concave base. The eyes were oblique, very long, banana-shaped, close-set and placed anteriorly on the carapace. Axes converged and crossed a short distance in front of the eyes. The ocelli were placed well forward between the eyes and the epistoma was well developed. The legs increased in size gradually, with the first three having short opposable spines at the end of each j ...
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Rhenopteroidea
The Rhenopteridae are a family of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The family is the only family currently contained in the superfamily Rhenopteroidea, one of four superfamilies classified as part of the suborder Stylonurina. The family contains one of the earliest known eurypterids, '' Brachyopterus'', known from the Middle Ordovician (also the sister taxon to all other rhenopterids), and is the most primitive clade of stylonurine eurypterids. The last known members of the family went extinct during the Early Devonian. Description Rhenopterids were small, characterized by scattered tubercules and knobs on the outer surface of the exoskeleton. Their first two (or possibly three) pairs of walking legs had spines; the last two pairs were long and powerful, without spines. The prosoma (head) was subtrapezoidal, with arcuate compound eyes on parallel axes. The male genital appendages were short with two distal spines. The ...
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Rhenopteridae
The Rhenopteridae are a family of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The family is the only family currently contained in the superfamily Rhenopteroidea, one of four superfamilies classified as part of the suborder Stylonurina. The family contains one of the earliest known eurypterids, '' Brachyopterus'', known from the Middle Ordovician (also the sister taxon to all other rhenopterids), and is the most primitive clade of stylonurine eurypterids. The last known members of the family went extinct during the Early Devonian. Description Rhenopterids were small, characterized by scattered tubercules and knobs on the outer surface of the exoskeleton. Their first two (or possibly three) pairs of walking legs had spines; the last two pairs were long and powerful, without spines. The prosoma (head) was subtrapezoidal, with arcuate compound eyes on parallel axes. The male genital appendages were short with two distal spines. The ...
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Rhenopterus
''Rhenopterus'' is an extinct prehistoric eurypterid. Fossils of ''Rhenopterus'' have been recovered from deposits of Lower Devonian age in Germany.Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 16.0 http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf (PDF). Description Rhenopterids were small, characterized by scattered tubercules and knobs on the outer surface of the exoskeleton. Their first two (or possibly three) pairs of walking legs had spines; the last two pairs were long and powerful, without spines. The prosoma (head) was subtrapezoidal, with arcuate compound eyes on parallel axes. The male genital appendages were short with two distal spines. Species ''Rhenopterus'' contains three valid species, with other named species now seen as invalid or part of other genera. * ''Rhenopterus diensti'' Størmer, 1936 - Germany (Lower D ...
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List Of Eurypterids
This list of eurypterid genera is a comprehensive listing of all genera that have ever been included in the order Eurypterida, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful ('' nomen dubium''), or were not formally published ('' nomen nudum''), as well as junior synonyms of more established names and genera that are no longer considered eurypterids. The list currently includes 115 names out of which 74 are considered valid eurypterid genera. There are approximately 250 species of eurypterids recognized as valid. Naming conventions and terminology There is no "official" or "canonical" list of eurypterid genera. The closest thing is found contained in the regularly updated ''Summary list'' ''of fossil spiders and their relatives'' in the World Spider Catalog. The vast majority of the content of the list below, including the valid genera, preoccupied names, junior synonyms, taxonomical ...
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Stylonurina
Stylonurina is one of two suborders of eurypterids, a group of extinct arthropods commonly known as "sea scorpions". Members of the suborder are collectively and informally known as "stylonurine eurypterids" or "stylonurines". They are known from deposits primarily in Europe and North America, but also in Siberia. Compared to the other suborder, Eurypterina, the stylonurines were comparatively rare and retained their posterior prosomal appendages for walking. Despite their rarity, the stylonurines have the longest temporal range of the two suborders. The suborder contains some of the oldest known eurypterids, such as ''Brachyopterus'', from the Middle Ordovician as well as the youngest known eurypterids, from the Late Permian. They remained rare throughout the Ordovician and Silurian, though the radiation of the mycteropoids (a group of large sweep-feeding forms) in the Late Devonian and Carboniferous is the last major radiation of the eurypterids before their extinction in the Pe ...
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Carapace
A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron. Crustaceans In crustaceans, the carapace functions as a protective cover over the cephalothorax (i.e., the fused head and thorax, as distinct from the abdomen behind). Where it projects forward beyond the eyes, this projection is called a rostrum (anatomy), rostrum. The carapace is Calcification, calcified to varying degrees in different crustaceans. Zooplankton within the phylum Crustacea also have a carapace. These include Cladocera, ostracods, and Isopoda, isopods, but isopods only have a developed "cephalic shield" carapace covering the head. Arachnids In arachnids, the carapace is formed by the fusion of prosomal tergites into a single Plate (animal anatomy), plate which carries the e ...
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Scotland
Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the north and west, the North Sea to the northeast and east, and the Irish Sea to the south. It also contains more than 790 islands, principally in the archipelagos of the Hebrides and the Northern Isles. Most of the population, including the capital Edinburgh, is concentrated in the Central Belt—the plain between the Scottish Highlands and the Southern Uplands—in the Scottish Lowlands. Scotland is divided into 32 administrative subdivisions or local authorities, known as council areas. Glasgow City is the largest council area in terms of population, with Highland being the largest in terms of area. Limited self-governing power, covering matters such as education, social services and roads and transportation, is devolved from the Scott ...
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Silurian Eurypterids
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone durin ...
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Pseudofossil
Pseudofossils are inorganic objects, markings, or impressions that might be mistaken for fossils. Pseudofossils may be misleading, as some types of mineral deposits can mimic lifeforms by forming what appear to be highly detailed or organized structures. One common example is when manganese oxides crystallize with a characteristic treelike or dendritic pattern along a rock fracture. The formation of frost dendrites on a window is another common example of this crystal growth. Concretions are sometimes thought to be fossils, and occasionally one contains a fossil, but are generally not fossils themselves. Chert or flint nodules in limestone can often take forms that resemble fossils. Background Pyrite disks or spindles are sometimes mistaken for fossils of sand dollars or other forms (see marcasite). Cracks, bumps, gas bubbles, and such can be difficult to distinguish from true fossils. Specimens that cannot be attributed with certainty to either fossils or pseudofossils are treated ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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Metasoma
The metasoma is the posterior part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory system, and circulatory system, and the apical segments are typically modified to form genitalia. In a few of the most primitive insects (the Archaeognatha), the metasomal segments bear small, articulated appendages called "styli", which are often considered to be vestigial. There are also pre-apical appendages in most insect orders, called cerci, which may be multi-segmented and almost resembling a posterior pair of antennae; these may be variously modified, or lost entirely. Otherwise, most adult insects lack appendages on the metasoma, though many larval insects (e.g., caterpillars) have some form of appendages, such as prolegs or, in aquatic insects, gills. In apocritan Hymenoptera (wasps, bees and ants), the metasoma consists of the second abdominal ...
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Arachnoid Granulation
Arachnoid granulations (also arachnoid villi, and pacchionian granulations or bodies) are small protrusions of the arachnoid mater (the thin second layer covering the brain) into the outer membrane of the dura mater (the thick outer layer). They protrude into the dural venous sinuses of the brain, and allow cerebrospinal fluid (CSF) to exit the subarachnoid space and enter the blood stream. The largest granulations lie along the superior sagittal sinus, a large venous space running from front to back along the center of the head (on the inside of the skull). They are, however, present along other dural sinuses as well. Function Diffusion across the arachnoid granulations into the superior sagittal sinus returns CSF to the venous circulation. The arachnoid granulations act as one-way valves. Normally the pressure of the CSF is higher than that of the venous system, so CSF flows through the villi and granulations into the blood. If the pressure is reversed for some reason, fluid ...
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