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Bombus Bifarius
''Bombus bifarius'', the two-form bumblebee, is a species of eusocial bumblebee of the subgenus ''Pyrobombus''. ''B. bifarius'' inhabits mountainous regions of western North America, primarily the states of Colorado and Utah. Its common name refers to a historical artifact, in that it was believed that this species had a color polymorphism, with a red-tailed nominate form and a black-tailed ''nearcticus'' form, present in the species. This polymorphism has recently been shown to belong instead to a cryptic sister species, '' Bombus vancouverensis'', that occupies almost the entirety of the range of what was formerly classified as ''bifarius''; true ''bifarius'' only has a red form, so it is not "two-formed" at all.Ghisbain, G., J. D. Lozier, S. R. Rahman, B. D. Ezray, L. Tian et al. (2020) Substantial genetic divergence and lack of recent gene flow support cryptic speciation in a colour polymorphic bumble bee (''Bombus bifarius'') species complex. Syst. Entomol. 45: 635–652. d ...
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Ezra Townsend Cresson
Ezra Townsend Cresson, also Ezra Townsend senior (18 June 1838, in Byberry – 19 April 1926, in Swarthmore) was an American entomologist who specialised in the Hymenoptera order of insects. He wrote ''Synopsis of the families and genera of the Hymenoptera of America, north of Mexico'' Philadelphia: Paul C. Stockhausen, Entomological printer (1887) and many other works. His son Ezra Townsend, Jr. (1876–1948) was also an entomologist but a specialist in Diptera. Cresson also documented many new species including ''Nomada texana ''Nomada texana'' is a species of bee native to the southern and western United States and other parts of North America (including Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sover ...''. References * Essig, E. O. 1931 ''A History of Entomology''. -New York, Macmillan Company. *Mallis, A. 1971 ''American Entomologists''. Rutgers Univ. Press New Brunswick 343-348, Portr. *Osborn, H. 1937 ...
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Eusocial
Eusociality (from Greek εὖ ''eu'' "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generations within a colony of adults, and a division of labor into reproductive and non-reproductive groups. The division of labor creates specialized behavioral groups within an animal society which are sometimes referred to as 'castes'. Eusociality is distinguished from all other social systems because individuals of at least one caste usually lose the ability to perform at least one behavior characteristic of individuals in another caste. Eusocial colonies can be viewed as superorganisms. Eusociality exists in certain insects, crustaceans, and mammals. It is mostly observed and studied in the Hymenoptera (ants, bees, and wasps) and in Blattodea (termites). A colony has caste differences: queens and reproductive males take the roles of the so ...
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Colorado
Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains. Colorado is the eighth most extensive and 21st most populous U.S. state. The 2020 United States census enumerated the population of Colorado at 5,773,714, an increase of 14.80% since the 2010 United States census. The region has been inhabited by Native Americans and their ancestors for at least 13,500 years and possibly much longer. The eastern edge of the Rocky Mountains was a major migration route for early peoples who spread throughout the Americas. "''Colorado''" is the Spanish adjective meaning "ruddy", the color of the Fountain Formation outcroppings found up and down the Front Range of the Rocky Mountains. The Territory of Colorado was organized on February 28, 1861, and on August 1, 1876, U.S. President Ulyss ...
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Utah
Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its west by Nevada. Utah also touches a corner of New Mexico in the southeast. Of the fifty U.S. states, Utah is the 13th-largest by area; with a population over three million, it is the 30th-most-populous and 11th-least-densely populated. Urban development is mostly concentrated in two areas: the Wasatch Front in the north-central part of the state, which is home to roughly two-thirds of the population and includes the capital city, Salt Lake City; and Washington County in the southwest, with more than 180,000 residents. Most of the western half of Utah lies in the Great Basin. Utah has been inhabited for thousands of years by various indigenous groups such as the ancient Puebloans, Navajo and Ute. The Spanish were the first Europe ...
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Polymorphism (biology)
In biology, polymorphism is the occurrence of two or more clearly different morphs or forms, also referred to as alternative ''phenotypes'', in the population of a species. To be classified as such, morphs must occupy the same habitat at the same time and belong to a panmictic population (one with random mating). Ford E.B. 1965. ''Genetic polymorphism''. Faber & Faber, London. Put simply, polymorphism is when there are two or more possibilities of a trait on a gene. For example, there is more than one possible trait in terms of a jaguar's skin colouring; they can be light morph or dark morph. Due to having more than one possible variation for this gene, it is termed 'polymorphism'. However, if the jaguar has only one possible trait for that gene, it would be termed "monomorphic". For example, if there was only one possible skin colour that a jaguar could have, it would be termed monomorphic. The term polyphenism can be used to clarify that the different forms arise from the s ...
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Bombus Vancouverensis
''Bombus vancouverensis'' is a common species of eusocial bumblebee of the subgenus ''Pyrobombus''. ''B. vancouverensis'' inhabits mountainous regions of western North America, where it has long been considered as a synonym of ''Bombus bifarius'', and essentially all of the literature on ''bifarius'' refers instead to ''vancouverensis''. ''B. vancouverensis'' has been identified as one of the two species of bumblebee observed to use pheromones in kin recognition. The other is the frigid bumblebee, ''Bombus frigidus''. Taxonomy ''Bombus vancouverensis'' has two recognized subspecies: *''Bombus vancouverensis vancouverensis'' Cresson, 1878 - limited to British Columbia *''Bombus vancouverensis nearcticus'' Handlirsch, 1888 - widespread in the United States and Canada ''Bombus vancouverensis'' was first described by Ezra Townsend Cresson in the 1878 ''Proceedings of the Academy of Natural Sciences of Philadelphia''. It is a member of the order Hymenoptera and the family Apidae, wh ...
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Hymenoptera
Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Parasitoid wasp, parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis (biology), metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they mature. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek language, Ancient Greek wikt:πτερόν, πτερόν (''pteron'') for wing. The Ancient Greek wikt:ὑμήν, ὑμήν (''hymen'') for membrane provides a plausible etymology for the term bec ...
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Apidae
Apidae is the largest family within the superfamily Apoidea, containing at least 5700 species of bees. The family includes some of the most commonly seen bees, including bumblebees and honey bees, but also includes stingless bees (also used for honey production), carpenter bees, orchid bees, cuckoo bees, and a number of other less widely known groups. Taxonomy In addition to its historical classification (honey bees, bumble bees, stingless bees and orchid bees), the family Apidae presently includes all the genera formerly placed in the families Anthophoridae and Ctenoplectridae. Although the most visible members of Apidae are social, the vast majority of apid bees are solitary, including a number of cleptoparasitic species. The old family Apidae contained four tribes (Apinae: Apini, Euglossini and Bombinae: Bombini, Meliponini) which have been reclassified as tribes of the subfamily Apinae, along with all of the former tribes and subfamilies of Anthophoridae and the former f ...
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Thorax
The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the creature's body, each of which is in turn composed of multiple segments. The human thorax includes the thoracic cavity and the thoracic wall. It contains organs including the heart, lungs, and thymus gland, as well as muscles and various other internal structures. Many diseases may affect the chest, and one of the most common symptoms is chest pain. Etymology The word thorax comes from the Greek θώραξ ''thorax'' "breastplate, cuirass, corslet" via la, thorax. Plural: ''thoraces'' or ''thoraxes''. Human thorax Structure In humans and other hominids, the thorax is the chest region of the body between the neck and the abdomen, along with its internal organs and other contents. It is mostly protected and supported by the rib cage, spi ...
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Pollen
Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophytes during the process of their movement from the stamens to the pistil of flowering plants, or from the male cone to the female cone of gymnosperms. If pollen lands on a compatible pistil or female cone, it germinates, producing a pollen tube that transfers the sperm to the ovule containing the female gametophyte. Individual pollen grains are small enough to require magnification to see detail. The study of pollen is called palynology and is highly useful in paleoecology, paleontology, archaeology, and forensics. Pollen in plants is used for transferring haploid male genetic material from the anther of a single flower to the stigma of another in cross-pollination. In a case of self-pollination, this process takes place from the anth ...
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Arizona
Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Four Corners region with Utah to the north, Colorado to the northeast, and New Mexico to the east; its other neighboring states are Nevada to the northwest, California to the west and the Mexican states of Sonora and Baja California to the south and southwest. Arizona is the 48th state and last of the contiguous states to be admitted to the Union, achieving statehood on February 14, 1912. Historically part of the territory of in New Spain, it became part of independent Mexico in 1821. After being defeated in the Mexican–American War, Mexico ceded much of this territory to the United States in 1848. The southernmost portion of the state was acquired in 1853 through the Gadsden Purchase. Southern Arizona is known for its desert cl ...
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New Mexico
) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Keres, Zuni , Governor = , Lieutenant Governor = , Legislature = New Mexico Legislature , Upperhouse = Senate , Lowerhouse = House of Representatives , Judiciary = New Mexico Supreme Court , Senators = * * , Representative = * * * , postal_code = NM , TradAbbreviation = N.M., N.Mex. , area_rank = 5th , area_total_sq_mi = 121,591 , area_total_km2 = 314,915 , area_land_sq_mi = 121,298 , area_land_km2 = 314,161 , area_water_sq_mi = 292 , area_water_km2 = 757 , area_water_percent = 0.24 , population_as_of = 2020 , population_rank = 36th , 2010Pop = 2,117,522 , population_density_rank = 45th , 2000DensityUS = 17.2 , 2000Density = 6.62 , MedianHouseholdIncome = $51,945 , IncomeRank = 45th , AdmittanceOrder = ...
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