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Bolosauria
Procolophonomorpha is an order or clade containing most parareptiles. Many papers have applied various definitions to the name, though most of these definitions have since been considered synonymous with modern parareptile clades such as Ankyramorpha and Procolophonia. The current definition of Procolophonomorpha, as defined by Modesto, Scott, & Reisz (2009), is that of as a stem-based group containing ''Procolophon'' and all taxa more closely related to it than to ''Milleretta''. It constitutes a diverse assemblage that includes a number of lizard-like forms, as well as more diverse types such as the pareiasaurs. Lee 1995, 1996, 1997 argues that turtles evolved from pareiasaurs, but this view is no longer considered likely. Rieppel and deBraga 1996 and deBraga and Rieppel, 1997 argue that turtles evolved from sauropterygians, and there is both molecular and fossil (''Pappochelys'') evidence for the origin of turtles among diapsid reptiles. Classification The following cladog ...
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Bolosauria
Procolophonomorpha is an order or clade containing most parareptiles. Many papers have applied various definitions to the name, though most of these definitions have since been considered synonymous with modern parareptile clades such as Ankyramorpha and Procolophonia. The current definition of Procolophonomorpha, as defined by Modesto, Scott, & Reisz (2009), is that of as a stem-based group containing ''Procolophon'' and all taxa more closely related to it than to ''Milleretta''. It constitutes a diverse assemblage that includes a number of lizard-like forms, as well as more diverse types such as the pareiasaurs. Lee 1995, 1996, 1997 argues that turtles evolved from pareiasaurs, but this view is no longer considered likely. Rieppel and deBraga 1996 and deBraga and Rieppel, 1997 argue that turtles evolved from sauropterygians, and there is both molecular and fossil (''Pappochelys'') evidence for the origin of turtles among diapsid reptiles. Classification The following cladog ...
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Erpetonyx
''Erpetonyx'' is an extinct genus of bolosaurian parareptile from the Gzhelian stage of the Carboniferous period, with a single known species: ''Erpetonyx arsenaultorum''. It is known from a single articulated and mostly complete specimen from Prince Edward Island in Canada. Phylogenetics has predicted that parareptiles first evolved in the Carboniferous, parallel to eureptiles ("true reptiles"). However, ''Hylonomus'', the oldest eureptile known from fossil evidence, lived millions of years before parareptiles appeared in the fossil record. The discovery of ''Erpetonyx'' helped to shorten this gap between parareptile and eureptile fossils, as ''Erpetonyx'' lived in the Late Carboniferous and is one of the oldest known parareptiles (though ''Carbonodraco'' is now known to be older). However, it was not closely related to ancestral parareptiles, so its discovery also indicated that the initial diversification of parareptiles occurred earlier in the Carboniferous. ''Erpetonyx'' was ...
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Microleter
''Microleter'' is an extinct genus of basal procolophonomorph parareptiles which lived in Oklahoma during the Early Permian period. The type and only known species is ''Microleter mckinzieorum''. ''Microleter'' is one of several parareptile taxa described from the Richards Spur fissure fills, and can be characterized from its high tooth count, lacrimal/ narial contact, short postfrontal, and slit-like temporal emargination edged by the postorbital, jugal, squamosal, and quadratojugal. Contrary to '' Australothyris'', which had a similar phylogenetic position as a basal procolophonomorph, ''Microleter'' suggests that early parareptile evolution occurred in Laurasia and that multiple lineages developed openings or emarginations in the temporal region. Discovery The only known specimen of ''Microleter'' is a well-preserved skull and lower jaw designated as OMNH 71306, the holotype specimen. It was found at the Dolese Brothers limestone quarry near Richards Spur in Comanch ...
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Ankyramorpha
Ankyramorpha ("anchor forms") is an extinct clade of procolophonomorph parareptiles which lived between the early Cisuralian epoch (middle Sakmarian stage) to the latest Triassic period (latest Rhaetian stage) of Africa, Antarctica, Asia, Australia, Europe, North America and South America.Jalil, N. E., & Janvier, P. (2005). Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d’Argana, Maroc. Geodiversitas, 27(1), 35-132. This clade was named in a 1996 parareptile study by Michael deBraga and Robert R. Reisz. They provided the name Ankyramorpha for a newly recognized clade encompassing "the most recent common ancestor of Procolophonia and Lanthanosuchoidea and all its descendants", and this clade name sees continued use among modern parareptile studies. A similar name, Hallucicrania, was provided in an earlier 1995 study by Michael S. Y. Lee, who defined it as the node-based taxon formed by the most recent common ancestor of lanthanosuchids and " ...
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Sauropterygia
Sauropterygia ("lizard flippers") is an extinct taxon of diverse, aquatic reptiles that developed from terrestrial ancestors soon after the end-Permian extinction and flourished during the Triassic before all except for the Plesiosauria became extinct at the end of that period. The plesiosaurs would continue to diversify until the end of the Mesozoic. Sauropterygians are united by a radical adaptation of their pectoral girdle, adapted to support powerful flipper strokes. Some later sauropterygians, such as the pliosaurs, developed a similar mechanism in their pelvis. Uniquely among reptiles, sauropterygians moved their tail vertically like modern cetaceans and sirenians. Origins and evolution The earliest sauropterygians appeared about 247 million years ago (Ma), at the start of the Middle Triassic: the first definite sauropterygian with exact stratigraphic datum lies within the Spathian division of the Olenekian era in South China. Early examples were small (around 60 c ...
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Pappochelys
''Pappochelys'' (''παπποχέλυς'' [πάππος (grandfather) + χέλυς (turtle)] meaning "grandfather turtle" in Ancient Greek, Greek) is an extinct genus of diapsid reptile possibly related to turtles. The genus contains only one species, ''Pappochelys rosinae'', from the Middle Triassic of Germany, which was named by paleontologists and Hans-Dieter Sues in 2015 in paleontology, 2015. The discovery of ''Pappochelys'' provides strong support for the placement of turtles within Diapsida, a hypothesis that has long been suggested by molecular data, but never previously by the fossil record. It is morphologically intermediate between the definite stem-turtle ''Odontochelys'' from the Late Triassic of China and ''Eunotosaurus'', a reptile from the Middle Permian of South Africa. Description ''Pappochelys'' had a wide body, small skull, and a long tail that makes up about half of the total body length, which is up to . The skull is pointed with large eye sockets. Several ...
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Diapsid
Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ago during the late Carboniferous period. All diapsids other than the most primitive ones in the clade Araeoscelidia are sometimes placed into the clade Neodiapsida. The diapsids are extremely diverse, and include birds and all modern reptile groups, including turtles, which were historically thought to lie outside the group. Although some diapsids have lost either one hole (lizards), or both holes (snakes and turtles), or have a heavily restructured skull (modern birds), they are still classified as diapsids based on their ancestry. At least 17,084 species of diapsid animals are extant: 9,159 birds, and 7,925 snakes, lizards, tuatara, turtles, and crocodiles. Characteristics The name Diapsida means "two arches", and diapsids are tradition ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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Gzhelian
The Gzhelian ( ) is an age in the ICS geologic timescale or a stage in the stratigraphic column. It is the youngest stage of the Pennsylvanian, the youngest subsystem of the Carboniferous. The Gzhelian lasted from to Ma. It follows the Kasimovian age/stage and is followed by the Asselian age/stage, the oldest subdivision of the Permian system. The Gzhelian is more or less coeval with the Stephanian Stage of the regional stratigraphy of Europe. Name and definition The Gzhelian is named after the Russian village of Gzhel (russian: Гжель), nearby Ramenskoye, not far from Moscow. The name and type locality were defined by Sergei Nikitin (1850–1909) in 1890. The base of the Gzhelian is at the first appearance of the Fusulinida genera '' Daixina'', '' Jigulites'' and '' Rugosofusulina'', or at the first appearance of the conodont ''Streptognathodus zethus''. The top of the stage (the base of the Permian system) is at the first appearance of the conodont ''Streptognath ...
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Milleretta
''Milleretta'' is an extinct genus of millerettid parareptile from the Late Permian of South Africa. Fossils have been found in the Balfour Formation.Ruta, M., Cisneros, JC., Liebrecht, T., Tsuji, L. A. and Müller, J. 2011Amniotes through major biological crises: faunal turnover among Parareptiles and the end-Permian mass extinction ''Palaeontology'', 54: 1117–1137. doi:10.1111/j.1475-4983.2011.01051. ''Milleretta'' was a moderately sized, lizard-like animal, about in length. It was probably insectivorous. Its only known species is ''Milleretta rubidgei'', making ''Milleretta'' a monospecific genus.Gow, C.E. 1997A Note on the Postcranial Skeleton of Milleretta (Amniota: Parareptilia) 34,55–57 Discovery The name provided for this genus upon Robert Broom's original 1938 description was '' 'Millerina' '', but it was later renamed in 1947 when Broom discovered that the name '' 'Millerina' '' had already been used for a genus of fly. The new name, ''Milleretta,'' means 'Miller's ...
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Parareptilia
Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as ''Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by the ...
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Mesosaurus
''Mesosaurus'' (meaning "middle lizard") is an extinct genus of reptile from the Early Permian of southern Africa and South America. Along with it, the genera '' Brazilosaurus'' and ''Stereosternum'', it is a member of the family Mesosauridae and the order Mesosauria. ''Mesosaurus'' was long thought to have been one of the first marine reptiles, although new data suggests that at least those of Uruguay inhabited a hypersaline water body, rather than a typical marine environment. In any case, it had many adaptations to a fully aquatic lifestyle. It is usually considered to have been anapsid, although Friedrich von Huene considered it to be a synapsid, and this hypothesis has been revived recently. Discovery and naming The holotype of ''M. tenuidens'', MNHN 1865-77, is nicknamed the "Griqua ''Mesosaurus''" and it was found in a Griqua hut in South Africa, likely in Kimberley, Northern Cape around 1830 and was being used as a pot lid. The circumstances of its discovery and how i ...
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