Blechnales
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Blechnales
Aspleniineae is a suborder of ferns in the order Polypodiales. It is equivalent to the clade eupolypods II in earlier systems; it is also treated as a single very broadly defined family Aspleniaceae. The suborder generally corresponds with the order Blechnales as described by J. L. Reveal in 1993. Aspleniineae includes some important ferns, including ''Onoclea sensibilis'', the sensitive fern, which grows as a virtual weed throughout much of its temperate North American range, and ferns of the genus ''Thelypteris'', a genus that has shown remarkable speciation. It also includes one of the more common horticultural ferns, ''Matteuccia struthiopteris'', the ostrich fern. Taxonomy In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the group is treated as the suborder Aspleniinae, and divided into 11 families. Alternatively, it may be treated as a single, very broadly circumscribed family Aspleniaceae ''sensu lato ''Sensu'' is a Latin word meaning "in the sense o ...
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Onocleaceae
Onocleaceae is a small family of terrestrial ferns in the order Polypodiales. It is placed in the suborder Aspleniineae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family, along with Blechnaceae, may be placed in a very broadly defined family Aspleniaceae as the subfamily Blechnoideae. The family may contain from one to four genera, consisting of five species largely in north temperate climes. The four genera, '' Matteuccia'', ''Onoclea'', '' Onocleopsis'' and '' Pentarhizidium'', may be included under the single genus ''Onoclea''. Description Members of the family have the following characteristics, being distinguished by having strongly dimorphic fronds, with the fertile fronds different from the sterile fronds. The rhizomes are long- to short-creeping to ascending, and sometimes stoloniferous (''Matteuccia'' and ''Onocleopsis''). The leaves are strongly dimorphic and the petioles have two vascular bundles uniting distally into a gut ...
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Asplenium Adiantum-nigrum
''Asplenium adiantum-nigrum'' is a common species of fern known by the common name black spleenwort. It is found mostly in Africa, Europe, and Eurasia, but is also native to a few locales in Mexico and the United States. Description This spleenwort has thick, triangular leaf blades up to 10 centimeters long which are divided into several subdivided segments. It is borne on a reddish green petiole and the rachis is shiny and slightly hairy. The undersides of each leaf segment have one or more sori arranged in chains.Esser, Lora L. 1994''Asplenium adiantum-nigrum''.In: Fire Effects Information System, nline U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Taxonomy Linnaeus was the first to describe black spleenwort with the binomial ''Asplenium adiantum-nigrum'' in his '' Species Plantarum'' of 1753. A chloroplast phylogeny verified the allopolyploid origin of ''A. adiantum-nigrum'', with '' A. cu ...
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Circumscription (taxonomy)
In biological taxonomy, circumscription is the content of a taxon, that is, the delimitation of which subordinate taxa are parts of that taxon. If we determine that species X, Y, and Z belong in Genus A, and species T, U, V, and W belong in Genus B, those are our circumscriptions of those two genera. Another systematist might determine that T, U, V, W, X, Y, and Z all belong in genus A. Agreement on circumscriptions is not governed by the Codes of Zoological or Botanical Nomenclature, and must be reached by scientific consensus. A goal of biological taxonomy is to achieve a stable circumscription for every taxon. This goal conflicts, at times, with the goal of achieving a natural classification that reflects the evolutionary history of divergence of groups of organisms. Balancing these two goals is a work in progress, and the circumscriptions of many taxa that had been regarded as stable for decades are in upheaval in the light of rapid developments in molecular phylogenetics ...
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Athyriaceae
The Athyriaceae (ladyferns and allies) are a family of terrestrial ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae, and includes two genera. Alternatively, it may be treated as the subfamily Athyrioideae of a very broadly defined family Aspleniaceae. The family has with a cosmopolitan distribution. Description Species of the Athyriaceae are terrestrial or lithophytic, less commonly aquatic. They grow from various kinds of rhizome: short or long, creeping or erect, branched or not. The distribution and evolution of characters in the family is complex, and the genera have few constant features by which they can be identified. The sporangia have stalks two or three cells wide in the middle, and contain brown monolete spores. Taxonomy Earlier classifications The family was first created by Arthur H.G. Alston in 1956. It has had a varied history. In 2014, Christenhusz ...
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Blechnaceae
Blechnaceae is a family of ferns in the order Polypodiales, with a cosmopolitan distribution. Its status as a family and the number of genera included have both varied considerably. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family has 24 genera, and excludes genera placed in the separate family Onocleaceae. The family is divided into three subfamilies, including Blechnoideae s.s. Alternatively, the entire family may be treated as the subfamily Blechnoideae s.l. of a very broadly defined family Aspleniaceae, and include genera others place in Onocleaceae. Description Most are ground dwelling, some are climbers, such as ''Stenochlaena''. A characteristic feature of many species is that the young opening fronds are usually tinged with red. Taxonomy The family was created by Newman in 1844. In 2014, Christenhusz and Chase submerged it as subfamily Blechnoideae within the family Aspleniaceae and included Onocleaceae in it. The PPG I classifica ...
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Woodsiaceae
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treated as the subfamily Woodsioideae of a very broadly defined family Aspleniaceae ''Sensu, sensu lato''. Species of ''Woodsia'' are commonly known as cliff ferns. Taxonomy Woodsiaceae formerly included the members of the families Athyriaceae and Diplaziopsidaceae, but analysis has consistently shown that they should be treated as separate families. The following cladogram for the suborder Aspleniineae (as eupolypods II), based on Lehtonen (2011), and Rothfels & al. (2012), shows a likely phylogenetics, phylogenetic relationship between the Woodsiaceae and the other families of the Aspleniineae. Species There are about 40–50 species of the genus ''Woodsia''. , ''Plants of the World Online'' accepted the following species: *''Woodsia alpi ...
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Hemidictyaceae
''Hemidictyum'' is a genus of ferns with a single species, ''Hemidictyum marginatum'', commonly known as the marginated half net fern. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Hemidictyaceae. Alternatively, the family, along with Aspleniaceae '' sensu stricto'', may be placed in a much more broadly defined family Aspleniaceae as the subfamily Asplenioideae. Taxonomy The name ''Hemidictyum'' was derived from the terms ''hemi'' (half) and ''diktyon'' (net), from the veins being netted only half-way across the pinnules.''The Fern Manual, being a description of all the best stove, greenhouse and hardy ferns'' by contributors to the ''Journal of Horticulture'' p.69, London, 1863 Phylogenetic relationships Hemidictyaceae is considered to be a sister family to Aspleniaceae ''s.l.'', believed to have diverged during the Cretaceous period. The following cladogram for the suborder Aspleniineae (as eupolypods II), based on Lehto ...
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Desmophlebiaceae
''Desmophlebium'' is a genus of ferns. In the Pteridophyte Phylogeny Group classification of 2016, it is the only genus in the family Desmophlebiaceae. Other sources place it in a more widely defined Aspleniaceae. Taxonomy The family and genus were erected in 2016, following a molecular phylogenetic study. It was shown that the species previously known as ''Diplazium lechleri'' was sister to a clade formed by the families Hemidictyaceae and Aspleniaceae, considerably removed from the Athyriaceae where it had previously been classified. Accordingly, a new genus and family were created and ''Diplazium lechleri'' transferred to '' Desmophlebium lechleri''. Another species was added to the genus on the basis of its morphology. The generic name refers to the distinctive thickened vein running just inside the edge of a pinna (submarginal) connecting the ends of other veins. The authors derived it from the Greek , , meaning 'band', combined with , , meaning 'vein'. ( is the genitive o ...
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Diplaziopsidaceae
Diplaziopsidaceae is a family of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae, and includes two genera. Alternatively, it may be treated as the subfamily Diplaziopsidoideae of a very broadly defined family Aspleniaceae. They are described as medium-to-large ferns, which grow near streams in forested areas. Their rhizomes are thick and decumbent to erect. Species are found in east Asia, from China south to New Guinea and east into the Pacific. Taxonomy Maarten J. M. Christenhusz and Xuan-Chun Zhang originally described the family in 2011 by including three genera ''Diplaziopsis'', ''Hemidictyum'', and '' Homalosorus''. Later that year Samuli Lehtonen found ''Hemidictyum'' to be a sister to Aspleniaceae, so ''Hemidictyum'' was placed in its own family, Hemidictyaceae. Christenhusz and Mark W. Chase later included ''Hemidictyum'' in their subfamily Asplenioideae rather ...
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Rhachidosoraceae
''Rhachidosorus'' is a genus of ferns in the order Polypodiales. It is the only genus in the family Rhachidosoraceae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the genus may be placed in the subfamily Rhachidosoroideae of a more broadly defined family Aspleniaceae, the family placement used in '' Plants of the World Online'' . Species The genus was described by Ren Chang Ching in 1964, with about 7 species in eastern and southeastern Asia, including Japan, the Philippines, and Sumatra. , '' Plants of the World Online'' accepted the following species: *'' Rhachidosorus blotianus'' Ching – China, Vietnam *'' Rhachidosorus chrysocarpus'' (Alderw.) Ching *'' Rhachidosorus consimilis'' Ching – China (Guizhou, Sichuan, Yunnan) *'' Rhachidosorus mesosorus'' (Makino) Ching – China, Japan, South Korea *'' Rhachidosorus pulcher'' (Tagawa) Ching – Taiwan, Yunnan *'' Rhachidosorus siamensis'' S.Linds. *'' Rhachidosorus stramineus'' (Copel.) Chi ...
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Cystopteridaceae
Cystopteridaceae is a family of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae of the order Polypodiales, and includes three genera. Alternatively, it may be treated as the subfamily Cystopteridoideae of a very broadly defined family Aspleniaceae. Cystopteridaceae are small or medium-sized ferns in forests and crevices. They generally have thin laminae, and small, round, naked sori. Genera Three genera are accepted in the PPG I classification, and by the ''Checklist of Ferns and Lycophytes of the World'' . One hybrid genus also exists: *'' Acystopteris'' Nakai *''Cystopteris'' Bernh. *''Gymnocarpium ''Gymnocarpium'' is a small genus of ferns, called oak ferns. It was once placed with various other groups, including the dryopteroid ferns and the athyrioid ferns. Cladistic analysis has demonstrated that ''Gymnocarpium'' and ''Cystopteris'' ...'' Newman *'' ×Cystoc ...
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Sensu
''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular concept, but it also appears in expressions that indicate the convention or context of the usage. Common qualifiers ''Sensu'' is the ablative case of the noun ''sensus'', here meaning "sense". It is often accompanied by an adjective (in the same case). Three such phrases are: *''sensu stricto'' – "in the strict sense", abbreviation ''s.s.'' or ''s.str.''; *''sensu lato'' – "in the broad sense", abbreviation ''s.l.''; *''sensu amplo'' – "in a relaxed, generous (or 'ample') sense", a similar meaning to ''sensu lato''. Søren Kierkegaard uses the phrase ''sensu eminenti'' to mean "in the pre-eminent r most important or significantsense". When appropriate, comparative and superlative adjectives may also be used to convey the meaning ...
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