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Bear-dog
Amphicyonidae is an extinct family of terrestrial carnivorans belonging to the suborder Caniformia. They first appeared in North America in the middle Eocene (around 45 mya), spread to Europe by the late Eocene (35 mya), and appear in Asia, and Africa by the early Miocene (23 mya). They had largely disappeared worldwide by the late Miocene (8 mya), with the latest recorded species at the end of the Miocene in Pakistan. They were among the first carnivorans to evolve large body size. Later in their history, they came into competition with hesperocyonine and borophagine canids. As dogs evolved similar body sizes and cranial and dental adaptations, the rise of these groups may have led to their extinction. Amphicyonids are often colloquially referred to as "bear-dogs". Taxonomy The family was erected by Haeckel (1886) lso attributed to Trouessart (1885) Their exact position has long been disputed. Some early paleontologists defined them as members of the family Canidae, but th ...
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Carnivora
Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are formally referred to as carnivorans, and have evolved to specialize in eating flesh. The order is the fifth largest order of mammals, comprising at least 279 species. Carnivorans live on every major landmass and in a variety of habitats, ranging from the cold polar regions to the hyper-arid region of the Sahara Desert to the open seas. They come in a very large array of different body plans in contrasting shapes and sizes. Carnivora can be divided into two subclades: the cat-like Feliformia and the dog-like Caniformia, which are differentiated based on the structure of their ear bones and cranial features. The feliforms include families such as the felidae, cats, the hyenas, the mongooses and the viverridae, civets. The majority of felifor ...
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Amphicyoninae
Amphicyoninae is a subfamily of extinct bear-dogs, large terrestrial carnivores belonging to the suborder Caniformia and which inhabited North America, Eurasia, and Africa from ~37.2—2.6  Ma. Amphicyoninae existed for approximately ~. Amphicyoninae was named by Trouessart (1885). It was assigned to Canidae by Matthew (1902); to Ursidae by Ginsburg (1977); and to Amphicyonidae by Hunt (1998). Genera include: *''Agnotherium'', found in both Europe and N. Africa *''Amphicyon'', found in both Europe and N. America *''Cynelos'', synonyms include ''Absonodaphoenus'' (from Florida) and ''Hecubides'' (from Africa), endemic to N. America *'' Cynodictis'' *''Ischyrocyon'', ''Hadrocyon'' is a synonym, endemic to N. America *''Goupilictis'' *''Magericyon'' *''Pliocyon'', endemic to N. America *''Pseudocyon'', ''Amphicyonopsis'' is a synonym, endemic to Europe and N. America *''Ysengrinia'', found in both Europe and N. America Fossil distribution Specimens have been recovered from: ...
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Caniformia
Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs (wolves, foxes, etc.), bears, raccoons, and mustelids. The Pinnipedia (seals, walruses and sea lions) are also assigned to this group. The center of diversification for the Caniformia is North America and northern Eurasia. Caniformia stands in contrast to the other suborder of Carnivora, the Feliformia ("cat-like" carnivorans), the center of diversification of which was in Africa and southern Asia. Description Most members of this group have nonretractile claws (the fisher, marten, sea otter ( forepaws only), red panda, and ringtail, and some foxes have retractile or semi-retractile claws) and tend to be plantigrade (with the exception of the Canidae). Other traits that separate the Caniformia from the Feliformia is that caniforms have longer jaws and more teeth, with less specialized carnassial teeth. They also tend more towards omnivory and opportunistic feeding, ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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Hesperocyoninae
The extinct Hesperocyoninae are one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Borophaginae and extant Caninae. Taxonomic history Hesperocyoninae are basal canids that gave rise to the other two canid subfamilies, the Borophaginae and Caninae. This subfamily was endemic to North America, living from the Duchesnean stage of the Late Eocene through to the early Barstovian stage of the Miocene, lasting around 20 million years. It comprises 10 recognized genera and 26 recognized species. Four major lineages can be defined based on shared characteristics: *''Mesocyon''-''Enhydrocyon'' clade (includes ''Cynodesmus, Sunkahetanka, Philotrox'') *''Osbornodon'' clade *'' Paraenhydrocyon'' *'' Ectopocynus clade'' The genus ''Caedocyon'', which is only known from a single partial cranium that shows some ''Paraenhydrocyon'' affinities, probably represents another independent lineage. ''Hesperocyon'', which lacks the shared derived c ...
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Borophaginae
The extinct Borophaginae form one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Hesperocyoninae and extant Caninae. Borophaginae, called "bone-crushing dogs", were endemic to North America during the Oligocene to Pliocene and lived roughly 34—2.5 million years ago, existing for about . Origin The Borophaginae descended from the subfamily Hesperocyoninae. The earliest and most primitive borophagine is the genus ''Archaeocyon'', which is a small fox-sized animal mostly found in the fossil beds in western North America. The borophagines soon diversified into several major groups. They evolved to become considerably larger than their predecessors, and filled a wide range of niches in late Cenozoic North America, from small omnivores to powerful, bear-sized carnivores, such as ''Epicyon''. Species There are 66 identified borophagine species, including 18 new ones that range from the Orellan to Blancan ages. A phylogenetic anal ...
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Canidae
Canidae (; from Latin, ''canis'', "dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found within the canid family, which are the extinct Borophaginae and Hesperocyoninae, and the extant Caninae. The Caninae are known as canines, and include domestic dogs, wolves, coyotes, foxes, jackals and other extant and extinct species. Canids are found on all continents except Antarctica, having arrived independently or accompanied human beings over extended periods of time. Canids vary in size from the gray wolf to the fennec fox. The body forms of canids are similar, typically having long muzzles, upright ears, teeth adapted for cracking bones and slicing flesh, long legs, and bushy tails. They are mostly social animals, living together in family units or small groups and behaving cooperatively. Typically, only the dominant pair in a group bree ...
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Daphoenodon
''Daphoenodon'' is an extinct genus of terrestrial carnivore, which lived in the early Miocene and belonged to the family Amphicyonidae ("bear dogs") of the suborder Caniformia Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs (wolves, foxes, etc.), bears, raccoons, and mustelids. The Pinnipedia (seals, walruses and sea lions) are also assigned to this group. .... The species of ''Daphoenodon'' are characterized by limbs that are specialized in fore and aft movement, as well as a body alignment that results in a lengthened stride. Hunt, R.M. (2009). "Long-Legged Pursuit Carnivorans (Amphicyonidae, Daphoeninae) from the Early Miocene of North America". ''Bulletin of the American Museum of Natural History.'' 318: 1-95. Species ''D.'' ''falkenbachi'' was a larger species that was found in northern Goshen, southeastern Platte Counties, Wyoming, and Nebraska. A smaller species, ''D. skinneri'', was found in southern ...
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Bear
Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found on the continents of North America, South America, Europe, and Asia. Common characteristics of modern bears include large bodies with stocky legs, long snouts, small rounded ears, shaggy hair, plantigrade paws with five nonretractile claws, and short tails. While the polar bear is mostly carnivorous, and the giant panda feeds almost entirely on bamboo, the remaining six species are omnivorous with varied diets. With the exception of courting individuals and mothers with their young, bears are typically solitary animals. They may be diurnal or nocturnal and have an excellent sense of smell. Despite their heavy build and awkward gait, they are adept runners, cli ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Digitigrade
In terrestrial vertebrates, digitigrade () locomotion is walking or running on the toes (from the Latin ''digitus'', 'finger', and ''gradior'', 'walk'). A digitigrade animal is one that stands or walks with its toes (metatarsals) touching the ground, and the rest of its foot lifted. Digitigrades include walking birds (what many assume to be bird knees are actually ankles), cats, dogs, and many other mammals, but not plantigrades or unguligrades. Digitigrades generally move more quickly and quietly than other animals. There are anatomical differences between the limbs of plantigrades, like humans, and both unguligrade and digitigrade limbs. Digitigrade and unguligrade animals have relatively long carpals and tarsals, and the bones which would correspond to the human ankle are thus set much higher in the limb than in a human. In a digitigrade animal, this effectively lengthens the foot, so much so that what are often thought of as a digitigrade animal's "hands" and "feet" correspon ...
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