Ammonites
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Ammonites
Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) than they are to shelled nautiloids such as the living ''Nautilus'' species. The earliest ammonites appeared during the Devonian, with the last species vanishing during the Cretaceous–Paleogene extinction event. Ammonites are excellent index fossils, and linking the rock layer in which a particular species or genus is found to specific geologic time periods is often possible. Their fossil shells usually take the form of planispirals, although some helically spiraled and nonspiraled forms (known as heteromorphs) have been found. The name "ammonite", from which the scientific term is derived, was inspired by the spiral shape of their fossilized shells, which somewhat resemble tightly coiled rams' horns. Pliny the Elder ( 79 AD near Pomp ...
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Heteromorph
The Ancyloceratina were a diverse suborder of ammonite most closely related to the ammonites of order Lytoceratina. They evolved during the Late Jurassic but were not very common until the Cretaceous period, when they rapidly diversified and became one of the most distinctive components of Cretaceous marine faunas. They have been recorded from every continent and many are used as zonal or index fossils. The most distinctive feature of the majority of the Ancyloceratina is the tendency for most of them to have shells that are not regular spirals like most other ammonites. These irregularly-coiled ammonites are called heteromorph ammonites, in contrast to regularly coiled ammonites, which are called homomorph ammonites. Biology The biology of the heteromorph ammonites is not clear, but one certainty is that their uncoiled shells would have made these forms very poor swimmers. Open shells, particularly ones with spines and ribs, create a lot of drag; but more importantly, the orien ...
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Cretaceous–Paleogene Extinction Event
The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the exception of some ectothermic species such as sea turtles and crocodilians, no tetrapods weighing more than survived. It marked the end of the Cretaceous Period, and with it the Mesozoic era, while heralding the beginning of the Cenozoic era, which continues to this day. In the geologic record, the K–Pg event is marked by a thin layer of sediment called the K–Pg boundary, which can be found throughout the world in marine and terrestrial rocks. The boundary clay shows unusually high levels of the metal iridium, which is more common in asteroids than in the Earth's crust. As originally proposed in 1980 by a team of scientists led by Luis Alvarez and his son Walter, it is now generally thought that the K–Pg extinction was cause ...
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Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ident ...
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Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating ...
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Goniatitida
Goniatids, informally goniatites, are Ammonoidea, ammonoid cephalopods that form the order Goniatitida, derived from the more primitive Agoniatitida during the Middle Devonian some 390 million years ago (around Eifelian stage). Goniatites (goniatitids) survived the Late Devonian extinction to flourish during the Carboniferous and Permian only to become Permian–Triassic extinction event, extinct at the end of the Permian some 139 million years later. Morphology All goniatites possessed an external shell, which is divided internally into chambers filled with gas giving it buoyancy during the life of the animal. An open chamber at the front of the shell provided living space for the goniatitid animal, with access to open water through a ventral siphuncle. The general morphology and habit of goniatites was probably similar to that of their later relatives the Ammonitida, ammonites, being free swimming and possessing a head with two well developed eyes and arms (or tentacles). G ...
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Nautiloid
Nautiloids are a group of marine cephalopods ( Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rel ...
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Mollusc
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8 taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropods ...
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Index Fossils
Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of Biology'', 8th ed., Oxford University Press, 2019. The primary objective of biostratigraphy is ''correlation'', demonstrating that a particular horizon in one geological section represents the same period of time as another horizon at a different section. Fossils within these strata are useful because sediments of the same age can look completely different, due to local variations in the sedimentary environment. For example, one section might have been made up of clays and marls, while another has more chalky limestones. However, if the fossil species recorded are similar, the two sediments are likely to have been laid down around the same time. Ideally these fossils are used to help identify biozones, as they make up the basic biostratigrap ...
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Coleoid
Subclass (biology), Subclass Coleoidea, or Dibranchiata, is the grouping of cephalopods containing all the various taxa popularly thought of as "soft-bodied" or "shell-less" (i.e., octopuses, squid and cuttlefish). Unlike its extant sister group, Nautiloidea, whose members have a rigid outer shell for protection, the coleoids have at most an internal cuttlebone, gladius (cephalopod), gladius, or shell that is used for buoyancy or support. Some species have lost their cuttlebone altogether, while in some it has been replaced by a chitinous support structure. A unique trait of the group is the ability to edit their own RNA. The major divisions of Coleoidea are based upon the number of cephalopod limb, arms or tentacles and their structure. The extinct and most primitive form, the Belemnoidea, presumably had ten equally-sized arms in five pairs numbered Dorsum (biology), dorsal to ventral as I, II, III, IV and V. More modern species either modified or lost a pair of arms. The superor ...
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Nautilus
The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina. It comprises six living species in two genera, the type of which is the genus ''Nautilus''. Though it more specifically refers to species ''Nautilus pompilius'', the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between 4 and 10 inches. Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle.Kümmel, B. 1964. Nautiloidae-Nautilida, in the Treatise on Invertebrate Paleontology, Geological Society of America and Univ of Kansas Press, Teichert and Moore eds. Having survived relatively u ...
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Clymeniida
Clymeniida is an order of ammonoid cephalopods from the Upper Devonian characterized by having an unusual dorsal siphuncle. They measured about in diameter and are restricted to Europe, North Africa, and possibly Australia. Morphologic characteristics Clymeniids produced a variety of shells ranging from smooth to ribbed and spinose and from evolute with all whorls exposed to strongly involute with the last whorl covering the previous. Some were even triangular as viewed from the side (along the axis of coiling). With the exception of the first few chambers, all have a siphuncle that runs along the dorsal margin, along the inside of the coils, opposite that of most ammonoids. The siphuncle starts off ventrally, like that in other ammonoids, but after the first few septa migrates to a definite and stable dorsal position. Septal necks are retrosiphonate, characteristic of their nautiloid ancestors, and are commonly very long, forming an almost continuous siphuncular tube. Sept ...
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Ceratitida
Ceratitida is an order that contains almost all ammonoid cephalopod genera from the Triassic as well as ancestral forms from the Upper Permian, the exception being the phylloceratids which gave rise to the great diversity of post Triassic ammonites. Ceratitids overwhelmingly produced planospirally coiled discoidal shells that may be evolute with inner whorls exposed or involute with only the outer whorl showing. In a few later forms the shell became subglobular, in others, trochoidal or uncoiled. Sutures are typically ceratitic, with smooth saddles and serrate or digitized lobes. In a few the sutures are goniatitic while in others they are ammonitic. Taxonomy * Ceratitida **Ceratitoidea ** Choristoceratoidea **Clydonitoidea **Danubitoidea **Dinaritoidea ** Lobitoidea ** Meekoceratoidea ** Megaphyllitoidea ** Nathorstitoidea **Noritoidea ** Otoceratoidea ** Pinacoceratoidea **Ptychitoidea ** Sageceratoidea **Tropitoidea ** Xenodiscoidea Only eight superfamilies are sho ...
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