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Alioramini
Alioramini is a tribe of long-snouted tyrannosaurine tyrannosaurids from the Late Cretaceous epoch. It includes the tyrannosaurid genera ''Alioramus'' and '' Qianzhousaurus''. Although tyrannosaurids are known from a variety of places around the globe, alioramins are currently restricted to Asia in mostly Maastrichtian strata. Alioramins were medium-sized tyrannosaurids, reaching around in length. Their snouts were much shallower and gracile than other tyrannosaurids, such as the massive ''Tarbosaurus'' or ''Tyrannosaurus''. Most notably, their nasal bones were ornamented with a series of rugose, pointy bumps on its top surface. The tribe Alioramini was first coined in 1995 by George Olshevsky only to contain the at-the-time uncertain ''Alioramus''. In 2014 the tribe was formally defined as a clade-based branch containing all tyrannosaurids more related to ''Alioramus'' than to ''Albertosaurus'', ''Proceratosaurus'', and ''Tyrannosaurus''. Hence, the tribe Alioramini consists of ...
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Qianzhousaurus
''Qianzhousaurus'' (meaning "Qianzhou lizard") is a genus of tyrannosaurid dinosaurs that lived in Asia during the Late Cretaceous period. There is currently only one species named, the type species ''Qianzhousaurus sinensis'' which is a member of the tribe Alioramini within the group, and most closely related to ''Alioramus'', another alioramin. History of discovery The holotype specimen, GM F10004, was unearthed in southern China, Ganzhou, at the Nanxiong Formation in the summer of 2010 during the construction of an industrial park and it was first described by paleontologists Junchang Lü, Laiping Yi, Stephen L. Brusatte, Ling Yang, Hua Li and Liu Chen in the journal ''Nature Communications'' in 2014. The genus is known from a partial sub-adult individual consisting of a nearly complete skull with the lower jaws missing all teeth (lost during fossilization), 9 cervical vertebrae, 3 dorsal vertebrae, 18 caudal vertebrae, both scapulocoracoids, partial ilia, and the left ...
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Alioramini Scale
Alioramini is a tribe of long-snouted tyrannosaurine tyrannosaurids from the Late Cretaceous epoch. It includes the tyrannosaurid genera '' Alioramus'' and ''Qianzhousaurus''. Although tyrannosaurids are known from a variety of places around the globe, alioramins are currently restricted to Asia in mostly Maastrichtian strata. Alioramins were medium-sized tyrannosaurids, reaching around in length. Their snouts were much shallower and gracile than other tyrannosaurids, such as the massive '' Tarbosaurus'' or ''Tyrannosaurus''. Most notably, their nasal bones were ornamented with a series of rugose, pointy bumps on its top surface. The tribe Alioramini was first coined in 1995 by George Olshevsky only to contain the at-the-time uncertain ''Alioramus''. In 2014 the tribe was formally defined as a clade-based branch containing all tyrannosaurids more related to ''Alioramus'' than to '' Albertosaurus'', ''Proceratosaurus'', and ''Tyrannosaurus''. Hence, the tribe Alioramini consists ...
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Tyrannosaurids
Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family (biology), family of coelurosaurian Theropoda, theropod dinosaurs that comprises two subfamilies containing up to thirteen genus, genera, including the eponymous ''Tyrannosaurus''. The exact number of genera is controversial, with some experts recognizing as few as three. All of these animals lived near the end of the Cretaceous Period (geology), Period and their fossils have been found only in North America and Asia. Although descended from Tyrannosauroidea, smaller ancestors, tyrannosaurids were almost always the largest Predation, predators in their respective ecosystems, putting them at the apex predator, apex of the food chain. The largest species was ''Tyrannosaurus rex'', one of the largest and most massive known land predators, which measured over in length and according to most modern estimates to in weight. Tyrannosaurids were Bipedalism, bipedal carnivores with massive skulls filled with large ...
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Tyrannosaurids
Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family (biology), family of coelurosaurian Theropoda, theropod dinosaurs that comprises two subfamilies containing up to thirteen genus, genera, including the eponymous ''Tyrannosaurus''. The exact number of genera is controversial, with some experts recognizing as few as three. All of these animals lived near the end of the Cretaceous Period (geology), Period and their fossils have been found only in North America and Asia. Although descended from Tyrannosauroidea, smaller ancestors, tyrannosaurids were almost always the largest Predation, predators in their respective ecosystems, putting them at the apex predator, apex of the food chain. The largest species was ''Tyrannosaurus rex'', one of the largest and most massive known land predators, which measured over in length and according to most modern estimates to in weight. Tyrannosaurids were Bipedalism, bipedal carnivores with massive skulls filled with large ...
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Alioramus
''Alioramus'' (; meaning 'different branch') is a genus of tyrannosaurid theropod dinosaurs from the Late Cretaceous period of Asia. It currently contains two species. The type species, ''A. remotus'' is known from a partial skull and three foot bones recovered from the Mongolian Nemegt Formation, which was deposited in a humid floodplain about 70 million years ago. These remains were named and described by Soviet paleontologist Sergei Kurzanov in 1976. A second species, ''A. altai'', known from a much more complete skeleton also from the Nemegt Formation, was named and described by Stephen L. Brusatte and colleagues in 2009. Its relationships to other tyrannosaurid genera were at first unclear, with some evidence supporting a hypothesis that ''Alioramus'' was closely related to the contemporary species ''Tarbosaurus bataar''. However, the discovery of ''Qianzhousaurus'' indicates that it belongs to a distinct branch of tyrannosaurs, namely the tribe Alioramini. ''Alioramu ...
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Alioramus Remotus
''Alioramus'' (; meaning 'different branch') is a genus of tyrannosaurid theropod dinosaurs from the Late Cretaceous period of Asia. It currently contains two species. The type species, ''A. remotus'' is known from a partial skull and three foot bones recovered from the Mongolian Nemegt Formation, which was deposited in a humid floodplain about 70 million years ago. These remains were named and described by Soviet paleontologist Sergei Kurzanov in 1976. A second species, ''A. altai'', known from a much more complete skeleton also from the Nemegt Formation, was named and described by Stephen L. Brusatte and colleagues in 2009. Its relationships to other tyrannosaurid genera were at first unclear, with some evidence supporting a hypothesis that ''Alioramus'' was closely related to the contemporary species ''Tarbosaurus bataar''. However, the discovery of '' Qianzhousaurus'' indicates that it belongs to a distinct branch of tyrannosaurs, namely the tribe Alioramini. ''Alior ...
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Tyrannosaurine
The tyrannosaurines (Tyrannosaurinae) is an extinct subfamily of Tyrannosauridae a family of coelurosaurian theropod dinosaurs that consists of at least three tribes and several genera. All fossils of these genera have been found in the Late Cretaceous deposits of western North America and east Asia. Compare to the related subfamily Albertosaurinae, tyrannosaurines overall are more robust and larger though the alioramins were gracile by comparison. This subfamily also includes the oldest known tyrannosaurid genus ''Lythronax'' as well as the youngest and most famous member of the group, ''Tyrannosaurus rex''. History of discovery The first remains of tyrannosaurids were uncovered during expeditions led by the Geological Survey of Canada, which located numerous scattered teeth. These distinctive dinosaur teeth were given the name ''Deinodon'' ("terrible tooth") by Joseph Leidy in 1856. In 1892 Edward Drinker Cope described more tyrannosaur material in the form of isolated vertebra ...
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Tarbosaurus
''Tarbosaurus'' ( ; meaning "alarming lizard") is a genus of tyrannosaurid dinosaur that flourished in Asia about 70 million years ago, at the end of the Late Cretaceous Period, considered to contain a single known species, ''Tarbosaurus bataar''. Fossils have been recovered from the Nemegt Formation of Mongolia, with more fragmentary remains found further afield in the Subashi Formation of China. Although many species have been named, modern paleontologists recognize only one, ''T. bataar'', as valid. Some experts see this species as an Asian representative of the North American genus ''Tyrannosaurus''; this would make the genus ''Tarbosaurus'' redundant. ''Tarbosaurus'' and ''Tyrannosaurus'', if not synonymous, are considered to be at least closely related genera. ''Alioramus'', also from Mongolia, has previously been thought by some authorities to be the closest relative of ''Tarbosaurus'', though this has since been disproven with the discovery of '' Qianzhousaurus'' and ...
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Tyrannosaurs
Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent beginning in the Jurassic Period. By the end of the Cretaceous Period, tyrannosauroids were the dominant large predators in the Northern Hemisphere, culminating in the gigantic ''Tyrannosaurus''. Fossils of tyrannosauroids have been recovered on what are now the continents of North America, Europe and Asia, with fragmentary remains possibly attributable to tyrannosaurs also known from South America and Australia. Tyrannosauroids were bipedal carnivores, as were most theropods, and were characterized by numerous skeletal features, especially of the skull and pelvis. Early in their existence, tyrannosauroids were small predators with long, three-fingered forelimbs. Late Cretaceous genera became much larger, including some of the largest land ...
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Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area of , about 30% of Earth's total land area and 8.7% of Earth's total surface area. The continent, which has long been home to the majority of the human population, was the site of many of the first civilizations. Its 4.7 billion people constitute roughly 60% of the world's population. In general terms, Asia is bounded on the east by the Pacific Ocean, on the south by the Indian Ocean, and on the north by the Arctic Ocean. The border of Asia with Europe is a historical and cultural construct, as there is no clear physical and geographical separation between them. It is somewhat arbitrary and has moved since its first conception in classical antiquity. The division of Eurasia into two continents reflects East–West cultural, linguistic, ...
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Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary extinction event) occurred at the end of this age. In this mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous. Stratigraphic definitions Definition The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying rock strata of the Chalk Group c ...
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Tribe (biology)
In biology, a tribe is a taxonomic rank above genus, but below family and subfamily. It is sometimes subdivided into subtribes. By convention, all taxonomic ranks from genus upwards are capitalized, including both tribe and subtribe. In zoology, the standard ending for the name of a zoological tribe is "-ini". Examples include the tribes Caprini (goat-antelopes), Hominini (hominins), Bombini (bumblebees), and Thunnini (tunas). The tribe Hominini is divided into subtribes by some scientists; subtribe Hominina then comprises "humans". The standard ending for the name of a zoological subtribe is "-ina". In botany, the standard ending for the name of a botanical tribe is "-eae". Examples include the tribes Acalypheae and Hyacintheae. The tribe Hyacintheae is divided into subtribes, including the subtribe Massoniinae. The standard ending for the name of a botanical subtribe is "-inae". In bacteriology, the form of tribe names is as in botany, e.g., Pseudomonadeae, based on the ge ...
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