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Acalyptratae
The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, as opposed to the Calyptratae. All forms of the name refer to the lack of calypters in the members of this subsection of flies. An alternative name, Acalypterae is current, though in minority usage. It was first used by Pierre-Justin-Marie Macquart in 1835 for a section of his tribe Muscides; he used it to refer to all acalyptrates plus scathophagids and phorids, but excluding Conopidae. The confusing forms of the names stem from their first usage; ''Acalyptratae'' and ''Acalyptrata'' actually are adjectival forms in New Latin. They were coined in the mid 19th century in contexts such as "Muscae Calyptratae and Acalyptratae" and "Diptera Acalyptrata", and the forms stuck. The Acalyptratae are a large assemblage, exhibiting very diverse h ...
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Sciomyzidae
The family Sciomyzidae belongs to the typical flies (Brachycera) of the order Diptera. They are commonly called marsh flies, and in some cases snail-killing flies due to the food of their larvae. Here, the Huttoninidae, Phaeomyiidae and Tetanoceridae are provisionally included in the Sciomyzidae. Particularly the latter seem to be an unequivocal part of this group and are ranked as tribe of subfamily Sciomyzinae by most modern authors, while the former two are very small lineages that may or may not stand outside the family and are provisionally ranked as subfamilies here. Whether the Salticellinae and the group around ''Sepedon'' warrant recognition as additional subfamilies or are better included in the Sciomyzinae proper is likewise not yet entirely clear. Altogether, the main point of contention is the relationship between the "Huttoninidae", "Phaeomyiidae", Sciomyzidae '' sensu stricto'', and the Helosciomyzidae which were also once included in the Sciomyzidae. Sciomy ...
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Ctenostylidae
The enigmatic fly family Ctenostylidae is a small group of very rare flies formerly included in the family Pyrgotidae (as the subfamily "Lochmostyliinae"); the principal reason for their inclusion in the Pyrgotidae was the absence of ocelli, a feature originally thought to be a unique defining feature ("autapomorphy") of the Pyrgotidae. Subsequent careful analysis has revealed that this anatomical feature shared with Pyrgotidae may not be indicative of a close relationship, and even the inclusion of Ctenostylidae within the superfamily Tephritoidea was cast into doubt, leaving this as the only family of Acalyptratae presently unassignable to superfamily.McAlpine, D.K. (1989) The taxonomic position of the Ctenostylidae (= Lochmostyliinae; Diptera: Schizophora). Mem. Inst. Oswaldo Cruz 84: 365-371 There are fewer than 20 described species in 7 genera in this pantropical family, all characterized not only by the lack of ocelli, but the lack of functional mouthparts, and unusual modifi ...
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Schizophora
The Schizophora are a section of true flies containing 78 families, which are collectively referred to as muscoids, although technically the term "muscoid" should be limited to flies in the superfamily Muscoidea; this is an example of informal, historical usage persisting in the vernacular. The section is divided into two subsections, the Acalyptratae and Calyptratae, which are commonly referred to as acalyptrate muscoids and calyptrate muscoids, respectively. The defining feature of the Schizophora is the presence of a special structure used to help the emerging adult fly break free of the puparium; this structure is an inflatable membranous sac called the ptilinum that protrudes from the face, above the antennae. The inflation of the ptilinum (using fluid hemolymph rather than air) creates pressure along the line of weakness in the puparium, which then bursts open along the seam to allow the adult to escape. When the adult emerges, the fluid is withdrawn, the ptilinum collapses ...
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Platystomatidae
The Platystomatidae (signal flies) are a distinctive family of flies (Diptera) in the superfamily Tephritoidea. Signal flies are worldwide in distribution, found in all the biogeographic realms, but predominate in the tropics. It is one of the larger families of acalyptrate Diptera with around 1200 species in 127 genera. Biology Adults are found on tree trunks and foliage and are attracted to flowers, decaying fruit, excrement, sweat, and decomposing snails. Larvae are found on fresh and decaying vegetation, carrion, human corpses, and root nodules, particularly in the genus ''Rivellia'', which has economic implications for legume crops. Larvae from the remaining genera are either phytophagous (eating plant material) or saprophagous (eating decomposing organic matter). Some are predatory on other insects and others have been found in human lesions, while others are of minor agricultural significance. Family description For terms see Morphology of Diptera Signal flies are very ...
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Lonchaeidae
The Lonchaeidae are a family of acalyptrate flies commonly known as lance flies. About 500 described species are placed into 9 genera. These are generally small but robustly built flies with blue-black or metallic bodies. They are found, mainly in wooded areas, throughout the world with the exception of polar regions and New Zealand. Description For terms see Morphology of Diptera. The Lonchaeids are small flies with a black or blackish-blue body, which is often metallic, glossy, and with hyaline (lacking dark spots) wings. The head is hemispherical (shorter than high) and the lunule is well defined. The third antennal segment is usually elongated and the antennae are decumbent. Ocelli are present and the postocellar bristles are divergent. The frons is narrow in males; in females it is broad. One pair of orbital bristles is on the head. The postvertical bristles on the head are parallel or weakly divergent. Distinctive vibrissae on the head are lacking, but vibrissa-like brist ...
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Pyrgotidae
The Pyrgotidae are an unusual family of flies (Diptera), one of only two families of Cyclorrhapha that lack ocelli. Most species are "picture-winged" (i.e., have patterns of bands or spots on the wings), as is typical among the Tephritoidea, but unlike other tephritoids, they are endoparasitoids; the females pursue scarab beetles in flight, laying an egg on the beetle's back under the elytra where the beetle cannot reach it. The egg hatches and the fly larva enters the body cavity of the beetle, feeding and eventually killing the host before pupating. In the United States, some species of '' Pyrgota'' and '' Sphecomyiella'' can be quite common in areas where their host beetles (typically the genus ''Phyllophaga'', or "June beetles") are abundant. Like their host beetles, these flies are primarily nocturnal, and are often attracted to artificial lights. Genera *''Acropyrgota'' Hendel, 1914 *'' Adapsilia'' Waga, 1842 *'' Adapsona'' Paramonov, 1958 *'' Afropyrgota'' V. Korneyev, ...
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Carnoidea
Carnoidea is a superfamily of Acalyptratae flies. Description In general, carnoids are small flies no more than a few millimetres long. Carnoidea is a poorly defined superfamily. In 1989, ten synapomorphies were described for the group, but most of these have later been challenged. As of 2006, the following synapopmorphies were described: uppermost fronto-orbital bristle(s) of the head is exclinate; phallus of the male is flexible, unsclerotized, simple and elongate; and phallus is microtrichose. Ecology Braulidae are associated with honey bees, with larvae developing in beeswax while adults attach to bees and feed from bee mouthparts. Canacidae adults are mainly found on seashore habitats such as beaches, estuarine tidal flats, wave-swept rocks and mangroves. Little is known about their larvae, but they are believed to mainly feed on algae in the intertidal zone. Australimyzidae are also found on seashores, being associated with dead or decaying plant matter. Carnidae are ...
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Calypter
A calypter is either of two posterior lobes of the posterior margin of the forewing of flies between the extreme posterior wing base and the alula, which covers the halteres. The lower calypter is the proximal calypter (synonyms: squama (of some authors), tegula) and the upper calypter is the distal calypter (synonym: squamula). Species of the subsection Acalyptratae The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, ... are noted for lacking calypters. References * James Francis McAlpine. ''Morphology and Terminology - Adults''. In: J.F. McAlpine, B.V. Peterson, G.E. Shewell, H.J. Teskey, J.R. Vockeroth, D.M. Wood (a cura di) ''Manual of Nearctic Diptera'', Volume 1. Research Branch, Agriculture Canada, Monograph 27, 1981: 9-64. . (In inglese). * Bernhard Merz, Jean-Paul Haenni. ...
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Diptera
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies and others, although only about 125,000 species have been described. Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great maneuverability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the l ...
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Ulidiidae
The Ulidiidae (formerly Otitidae) or picture-winged flies are a large and diverse cosmopolitan family of flies (Diptera), and as in related families, most species are herbivorous or detritivorous. They are often known as picture-winged flies, along with members of other families in the superfamily Tephritoidea that have patterns of bands or spots on the wings. Some species share with the Tephritidae an unusual elongated posteroapical projection of the anal cell in the wing, but can be differentiated by the smoothly curving subcostal vein. Two species, '' Tetanops myopaeformis'' and '' Euxesta stigmatias'', are agricultural pests. Systematics The Ulidiidae are divided into two subfamilies. Subfamily Otitinae ;Tribe Cephaliini Schiner, 1864 :*'' Acrostictella'' Hendel, 1914 :*'' Cephalia'' Meigen, 1826 :*'' Delphinia'' Robineau-Desvoidy, 1830 :*''Myiomyrmica'' Steyskal, 1961 :*''Myrmecothea'' Hendel, 1910 :*'' Proteseia'' Korneyev & Hernandes, 1998 :*'' Pterotaenia'' R ...
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Sciomyzoidea
Sciomyzoidea is a superfamily of Acalyptratae flies. The families placed here include at least the following: * Coelopidae – seaweed flies * Dryomyzidae * Helcomyzidae * Helosciomyzidae * Heterocheilidae * Huttoninidae * Natalimyzidae * Phaeomyiidae * Ropalomeridae * Sciomyzidae – marsh flies, snail-killing flies (including Huttoninidae, Phaeomyiidae, Tetanoceridae) * Sepsidae The Sepsidae are a family of flies, commonly called the black scavenger flies or ensign flies. Over 300 species are described worldwide. They are usually found around dung or decaying plant and animal material. Many species resemble ants, having ... – scavenger flies References External links * Diptera superfamilies Taxa named by Carl Fredrik Fallén {{Sciomyzoidea-stub ...
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Nerioidea
Nerioidea is a superfamily of Acalyptratae flies. Description As flies, Nerioidea undergo complete metamorphosis with the four life stages of egg, larva, pupa and adult. The adult stage has three body segments (head, thorax and abdomen), three pairs of legs and one pair of wings. Some features that distinguish adult Nerioidea from other flies are: a face that's usually weakly sclerotised (except in Fergusoninidae), antenna usually porrect or slightly deflexed (elbowed in Tanypezidae), wing veins R2+3 and R4+5 usually convergent, and the wing anal cell usually much smaller than the subcostal cell. Ecology Most Nerioidea are associated with dead and decaying organic matter such as dead wood, rotting fruit and bat dung. On the other hand, Fergusoninidae form galls in plants of family Myrtaceae, and some Micropezidae The Micropezidae are a moderate-sized family of acalyptrate muscoid flies in the insect order Diptera, comprising about 500 species in about 50 genera and five ...
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