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The Pallas's cat (''Otocolobus manul''), also known as the manul, is a small wild cat with long and dense light grey fur, and rounded ears set low on the sides of the head. Its head-and-body length ranges from with a long bushy tail. It is well camouflaged and adapted to the cold
continental climate Continental climates often have a significant annual variation in temperature (warm to hot summers and cold winters). They tend to occur in central and eastern parts of the three northern-tier continents (North America, Europe, and Asia), typi ...
in its native range, which receives little rainfall and experiences a wide range of temperatures. The Pallas's cat was first described in 1776 by
Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussia, Prussian zoologist, botanist, Ethnography, ethnographer, Exploration, explorer, Geography, geographer, Geology, geologist, Natura ...
, who observed it in the vicinity of
Lake Baikal Lake Baikal is a rift lake and the deepest lake in the world. It is situated in southern Siberia, Russia between the Federal subjects of Russia, federal subjects of Irkutsk Oblast, Irkutsk Oblasts of Russia, Oblast to the northwest and the Repu ...
. Since then, it has been recorded across a large region in
Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...
, albeit in widely spaced sites from the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
,
Iranian Plateau The Iranian plateau or Persian plateau is a geological feature spanning parts of the Caucasus, Central Asia, South Asia, and West Asia. It makes up part of the Eurasian plate, and is wedged between the Arabian plate and the Indian plate. ...
,
Hindu Kush The Hindu Kush is an mountain range in Central Asia, Central and South Asia to the west of the Himalayas. It stretches from central and eastern Afghanistan into northwestern Pakistan and far southeastern Tajikistan. The range forms the wester ...
, parts of the
Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...
,
Tibetan Plateau The Tibetan Plateau, also known as the Qinghai–Tibet Plateau or Qingzang Plateau, is a vast elevated plateau located at the intersection of Central Asia, Central, South Asia, South, and East Asia. Geographically, it is located to the north of H ...
to the
Altai-Sayan region The Altai-Sayan region is an area of Inner Asia proximate to the Altai Mountains and the Sayan Mountains, near to where Russia, China, Mongolia and Kazakhstan come together. This region is one of the world centers of Temperate climate, temperate pl ...
and South Siberian Mountains. It inhabits rocky
montane grasslands and shrublands Montane grasslands and shrublands are a biome defined by the World Wildlife Fund. The biome includes high elevation grasslands and shrublands around the world. The term "montane" in the name of the biome refers to "high elevation", rather than t ...
, where the snow cover is below . It finds shelter in rock crevices and
burrow file:Chipmunk-burrow (exits).jpg, An eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of Animal lo ...
s, and preys foremost on lagomorphs and
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s. The female gives birth to between two and six kittens in spring. Due to its widespread range and assumed large population, the Pallas's cat has been listed as
Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
since 2020. Some population units are threatened by
poaching Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...
, prey base decline due to rodent control programs, and
habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological proces ...
as a result of
mining Mining is the Resource extraction, extraction of valuable geological materials and minerals from the surface of the Earth. Mining is required to obtain most materials that cannot be grown through agriculture, agricultural processes, or feasib ...
and infrastructure projects. The Pallas's cat has been kept in zoos since the early 1950s. 60 zoos in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
, Russia, North America and
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
participate in Pallas's cat
captive breeding Captive breeding, also known as captive propagation, is the process of keeping plants or animals in controlled environments, such as wildlife reserves, zoos, Botanical garden, botanic gardens, and other Conservation biology, conservation facilitie ...
programs.


Etymology

'Manul' is the Pallas's cat's name in the
Mongolian language Mongolian is the Prestige (sociolinguistics), principal language of the Mongolic languages, Mongolic language family that originated in the Mongolian Plateau. It is spoken by ethnic Mongols and other closely related Mongolic peoples who are nati ...
. It is called 'manol' in the
Kyrgyz language Kyrgyz is a Turkic language of the Kipchak branch spoken in Central Asia. Kyrgyz is the official language of Kyrgyzstan and a significant minority language in the Kizilsu Kyrgyz Autonomous Prefecture in Xinjiang, China and in the Gorno-Badak ...
. The
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
'Pallas's cat' was coined by
William Thomas Blanford William Thomas Blanford (7 October 183223 June 1905) was an England, English geologist and natural history, naturalist. He is best remembered as the editor of a major series on ''The Fauna of British India, Including Ceylon and Burma''. Biogra ...
in honour of
Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussia, Prussian zoologist, botanist, Ethnography, ethnographer, Exploration, explorer, Geography, geographer, Geology, geologist, Natura ...
.


Taxonomy

''Felis manul'' was the
scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
used by
Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussia, Prussian zoologist, botanist, Ethnography, ethnographer, Exploration, explorer, Geography, geographer, Geology, geologist, Natura ...
in 1776, who first described a Pallas's cat that he had encountered near the Dzhida River southeast of
Lake Baikal Lake Baikal is a rift lake and the deepest lake in the world. It is situated in southern Siberia, Russia between the Federal subjects of Russia, federal subjects of Irkutsk Oblast, Irkutsk Oblasts of Russia, Oblast to the northwest and the Repu ...
. Several Pallas's cat zoological specimens were subsequently described: *''Felis nigripectus'' proposed by
Brian Houghton Hodgson Brian Houghton Hodgson (1 February 1801 – 23 May 1894) was a pioneer natural history, naturalist and ethnologist working in India and Nepal where he was a British Resident (title), Resident. He described numerous species of birds and mammals fr ...
in 1842 was based on three specimens from
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
. *''Otocolobus manul ferrugineus'' proposed by Sergey Ognev in 1928 was an erythristic specimen from the Kopet Dag mountains. ''Otocolobus'' was proposed by Johann Friedrich von Brandt in 1842 as a generic name.
Reginald Innes Pocock Reginald Innes Pocock, (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward's ...
recognized the
taxonomic rank In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of or ...
of ''Otocolobus'' in 1907, described several Pallas's cat skulls in detail and considered the Pallas's cat an aberrant form of '' Felis''. In 1951, John Ellerman and Terence Morrison-Scott considered *the
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
''Felis manul manul'' to be distributed from
Russian Turkestan Russian Turkestan () was a colony of the Russian Empire, located in the western portion of the Central Asian region of Turkestan. Administered as a Krai or Governor-Generalship, it comprised the oasis region to the south of the Kazakh Steppe, b ...
to Transbaikalia; *''F. m. nigripecta'' to be distributed in Tibet and
Kashmir Kashmir ( or ) is the Northwestern Indian subcontinent, northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term ''Kashmir'' denoted only the Kashmir Valley between the Great Himalayas and the Pir P ...
; *''F. m. ferruginea'' occurring from southwestern Turkestan and the Kopet Dag mountains to
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan, is a landlocked country located at the crossroads of Central Asia and South Asia. It is bordered by Pakistan to the Durand Line, east and south, Iran to the Afghanistan–Iran borde ...
and
Balochistan Balochistan ( ; , ), also spelled as Baluchistan or Baluchestan, is a historical region in West and South Asia, located in the Iranian plateau's far southeast and bordering the Indian Plate and the Arabian Sea coastline. This arid region o ...
. Since 2017, the Cat Classification Task Force of the Cat Specialist Group recognises only two subspecies as valid
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
, namely: *''O. m. manul'' syn. ''O. m. ferrugineus'' in the western and northern part of
Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...
from
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
to
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
; *''O. m. nigripectus'' in the
Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...
from Kashmir to
Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...
.


Phylogeny

Phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analysis of the
nuclear DNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. ...
in tissue samples from all Felidae species revealed that the
evolutionary radiation An evolutionary radiation is an increase in taxonomic diversity that is caused by elevated rates of speciation, that may or may not be associated with an increase in morphological disparity. A significantly large and diverse radiation within ...
of the Felidae began in Asia during the
late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million ye ...
around . Analysis of
mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
of all Felidae species indicates a radiation at around . The Pallas's cat is estimated to have genetically diverged from a
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonl ...
with the genus ''
Prionailurus ''Prionailurus'' is a genus of spotted, small wild cats native to Asia. Forests are their preferred habitat; they feed on small mammals, reptiles and birds, and occasionally aquatic wildlife. Taxonomy ''Prionailurus'' was first proposed by ...
'' between based on analysis of nuclear DNA. Based on analysis of mitochondrial DNA, it diverged from a common ancestor with ''Felis''.


Characteristics

The Pallas's cat's
fur A fur is a soft, thick growth of hair that covers the skin of almost all mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an ...
is light grey with pale yellowish-
ochre Ochre ( ; , ), iron ochre, or ocher in American English, is a natural clay earth pigment, a mixture of ferric oxide and varying amounts of clay and sand. It ranges in colour from yellow to deep orange or brown. It is also the name of the colou ...
or pale yellowish-reddish hues. Some hair tips are white and some blackish. Its fur is greyer and denser with fewer markings visible in winter than in the summer. The forehead and top of the head are light grey with small black spots. It has two black zigzag lines on the cheeks running from the corner of the eyes to the jaw joints. Its chin, whiskers, lower and upper lips are white. It has narrow black stripes on the back, consisting of five to seven dark transversal lines across the lower back. Its grey tail has seven narrow black rings and a black tip. The underfur is long and 19 μm thick, and the
guard hair Guard hair or overhair is the outer layer of hair of most mammals, which overlay the fur. Guard hairs are long and coarse and protect the rest of the pelage (fur) from abrasion and frequently from moisture. They are visible on the surface of the ...
s up to long and 93 μm thick on the back. Its fur is soft and dense with up to . The Pallas's cat's ears are grey with a yellowish tinge on the back and a darker rim, but with whitish hair in front and in the ear pinnae. Its rounded ears are set low on the side, such that it can peer over an object and show only a relatively small part of the head above the eyes without depressing the ears. This can give its face a look of ferocity and unrest. Its eyes are encircled by white. The iris is yellowish, and its
pupil The pupil is a hole located in the center of the iris of the eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing Company. It appears black becau ...
s contract to small circular disks in sunlight. Among the
Felinae The small cats or Felinae are a subfamily of Felidae distinguished by their bony hyoids, which let them purr but not roar. Other authors have proposed an alternative definition for this subfamily, as comprising only the living conical-toothed cat ...
, it shares this trait of round pupils with '' Puma'', '' Herpailurus'' and ''
Acinonyx ''Acinonyx'' is a genus within the Felidae family. The only living species of the genus, the cheetah (''A. jubatus''), lives in open grasslands of Africa and Asia. Several fossil remains of cheetah-like cats were excavated that date to the lat ...
'' species. The Pallas's cat is about the size of a
domestic cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small Domestication, domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have sh ...
(''Felis catus''). Its stocky posture with the long and dense fur make it appear stout and plush. Its head-to-body is long with a long tail. It weighs . Its body is stout, and its
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
is rounded with a short
nasal bone The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face and by their junction, form the bridge of the upper one third of the nose. Eac ...
, an enlarged
cranial Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Thi ...
part and rounded
zygomatic arch In anatomy, the zygomatic arch (colloquially known as the cheek bone), is a part of the skull formed by the zygomatic process of temporal bone, zygomatic process of the temporal bone (a bone extending forward from the side of the skull, over the ...
es. Its
orbits In celestial mechanics, an orbit (also known as orbital revolution) is the curved trajectory of an physical body, object such as the trajectory of a planet around a star, or of a natural satellite around a planet, or of an satellite, artificia ...
are large and directed forward. Its legs are short with short and sharp retractile claws. The skull of males is long and wide at the base. Females have a long and wide skull. The lower
carnassial Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified f ...
teeth are powerful, and the upper carnassials are short and massive. The first pair of upper
premolar The premolars, also called premolar Tooth (human), teeth, or bicuspids, are transitional teeth located between the Canine tooth, canine and Molar (tooth), molar teeth. In humans, there are two premolars per dental terminology#Quadrant, quadrant in ...
s is absent. The dental formula is . It has a bite force at the canine tip of 155.4
newtons The newton (symbol: N) is the unit of force in the International System of Units (SI). Expressed in terms of SI base units, it is 1 kg⋅m/s2, the force that accelerates a mass of one kilogram at one metre per second squared. The unit i ...
and a bite force quotient at the canine tip of 113.8. The mitochondrial genome of the Pallas's cat consists of 16,672
base pairs A base pair (bp) is a fundamental unit of double-stranded nucleic acids consisting of two nucleobases bound to each other by hydrogen bonds. They form the building blocks of the DNA double helix and contribute to the folded structure of both DNA ...
containing 13 protein-coding, 22
transfer RNA Transfer ribonucleic acid (tRNA), formerly referred to as soluble ribonucleic acid (sRNA), is an adaptor molecule composed of RNA, typically 76 to 90 nucleotides in length (in eukaryotes). In a cell, it provides the physical link between the gene ...
and two
ribosomal RNA Ribosomal ribonucleic acid (rRNA) is a type of non-coding RNA which is the primary component of ribosomes, essential to all cells. rRNA is a ribozyme which carries out protein synthesis in ribosomes. Ribosomal RNA is transcribed from ribosomal ...
genes and one
non-coding RNA A non-coding RNA (ncRNA) is a functional RNA molecule that is not Translation (genetics), translated into a protein. The DNA sequence from which a functional non-coding RNA is transcribed is often called an RNA gene. Abundant and functionally imp ...
control region.


Distribution and habitat

The Pallas's cat's range extends from the Caucasus eastward to Central Asia, Mongolia and adjacent parts of Dzungaria and the
Tibetan Plateau The Tibetan Plateau, also known as the Qinghai–Tibet Plateau or Qingzang Plateau, is a vast elevated plateau located at the intersection of Central Asia, Central, South Asia, South, and East Asia. Geographically, it is located to the north of H ...
. It inhabits montane
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
s and
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s, rocky
outcrop An outcrop or rocky outcrop is a visible exposure of bedrock or ancient superficial deposits on the surface of the Earth and other terrestrial planets. Features Outcrops do not cover the majority of the Earth's land surface because in most p ...
s,
scree Scree is a collection of broken rock fragments at the base of a cliff or other steep rocky mass that has accumulated through periodic rockfall. Landforms associated with these materials are often called talus deposits. The term ''scree'' is ap ...
slope In mathematics, the slope or gradient of a Line (mathematics), line is a number that describes the direction (geometry), direction of the line on a plane (geometry), plane. Often denoted by the letter ''m'', slope is calculated as the ratio of t ...
s and ravines in areas, where the continuous snow cover is below . In the southwestern part of its range, the habitat of the Pallas's cat is affected by cold and dry winters, and moderate to low rainfall in warm summers. The typical vegetation in this part consists of small
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s,
sagebrush Sagebrush is the common name of several woody and herbaceous species of plants in the genus ''Artemisia (plant), Artemisia''. The best-known sagebrush is the shrub ''Artemisia tridentata''. Sagebrush is native to the western half of North Amer ...
(''Artemisia''), ''
Festuca ''Festuca'' (fescue) is a genus of flowering plants belonging to the grass family Poaceae (subfamily Pooideae). They are evergreen or herbaceous perennial tufted grasses with a height range of and a cosmopolitan distribution, occurring on ...
'' and ''
Stipa ''Stipa'' is a genus of 141 species of large Perennial plant, perennial hermaphrodite, hermaphroditic grasses collectively known as feather grass, needle grass, and spear grass. They are placed in the subfamily Pooideae and the tribe Stipeae, w ...
'' grasses. In the central part of its range, it inhabits hilly landscapes, high
plateau In geology and physical geography, a plateau (; ; : plateaus or plateaux), also called a high plain or a tableland, is an area of a highland consisting of flat terrain that is raised sharply above the surrounding area on at least one side. ...
s and intermontane valleys that are covered by dry steppe or semi-desert vegetation, such as low shrubs and
xerophytic A xerophyte () is a species of plant that has adaptations to survive in an environment with little liquid water. Examples of xerophytes include cactus, cacti, pineapple and some gymnosperm plants. The morphology (biology), morphology and physiology ...
grasses. The
continental climate Continental climates often have a significant annual variation in temperature (warm to hot summers and cold winters). They tend to occur in central and eastern parts of the three northern-tier continents (North America, Europe, and Asia), typi ...
in this region exhibits a range of between the highest and lowest air temperatures, dropping to in winter. The Greater Caucasus region is considered climatically suitable for the Pallas's cat. In
Armenia Armenia, officially the Republic of Armenia, is a landlocked country in the Armenian Highlands of West Asia. It is a part of the Caucasus region and is bordered by Turkey to the west, Georgia (country), Georgia to the north and Azerbaijan to ...
, an individual was killed near Vedi in the mountains of
Ararat Province Ararat (, ) is a administrative divisions of Armenia, province (''marz (territorial entity), marz'') of Armenia. Its capital and largest city is the town of Artashat, Armenia, Artashat. The province is named after the biblical Mount Ararat. It i ...
in the late 1920s. In January 2020, an individual was sighted about farther north in Tavush Province; the habitat at this location transitions from semi-desert to montane steppe at an elevation of about . Records in
Azerbaijan Azerbaijan, officially the Republic of Azerbaijan, is a Boundaries between the continents, transcontinental and landlocked country at the boundary of West Asia and Eastern Europe. It is a part of the South Caucasus region and is bounded by ...
are limited to a Pallas's cat skin found in
Karabakh Karabakh ( ; ) is a geographic region in southwestern Azerbaijan and eastern Armenia, extending from the highlands of the Lesser Caucasus down to the lowlands between the rivers Kura and Aras. It is divided into three regions: Highland Kara ...
and a sighting of an individual in
Julfa District Julfa District () is one of the 7 districts of the Nakhchivan Autonomous Republic of Azerbaijan. The district borders the districts of Ordubad, Babek, Shahbuz, as well as the Syunik Province of Armenia and the East Azerbaijan Province of Ira ...
, both in the late 20th century. On the Iranian Plateau, two Pallas's cats were encountered near the
Aras River The Aras is a transboundary river in the Caucasus. It rises in eastern Turkey and flows along the borders between Turkey and Armenia, between Turkey and the Nakhchivan exclave of Azerbaijan, between Iran and both Azerbaijan and Armenia, and, fin ...
in northwestern Iran before the 1970s. In the area, an individual was captured at an elevation of about near Azarshahr in
East Azerbaijan Province East Azerbaijan province () is one of the 31 provinces of Iran. Its capital is the city of Tabriz. The province is located in Azerbaijan (Iran), Iranian Azerbaijan, bordering Armenia, the Azerbaijan, Republic of Azerbaijan, Ardabil pro ...
in 2008. In the same year, a camera trap recorded a Pallas's cat on the southern slopes of the central
Alborz Mountains The Alborz ( ) range, also spelled as Alburz, Elburz or Elborz, is a mountain range in northern Iran that stretches from the border of Azerbaijan along the western and entire southern coast of the Caspian Sea and finally runs northeast and merge ...
in Khojir National Park shortly after heavy snowfall. Farther east in the Alborz Mountains, an individual was recorded among rocks at an elevation of in 2016. In the Aladagh and Kopet Dag Mountains, the Pallas's cat was recorded inside and in the vicinity of protected areas. In the south of the
Zagros Mountains The Zagros Mountains are a mountain range in Iran, northern Iraq, and southeastern Turkey. The mountain range has a total length of . The Zagros range begins in northwestern Iran and roughly follows Iran's western border while covering much of s ...
, an individual was caught in a corral used by
transhumant Transhumance is a type of pastoralism or nomadism, a seasonal movement of livestock between fixed summer and winter pastures. In montane regions (''vertical transhumance''), it implies movement between higher pastures in summer and lower ...
pastoralists in Abadeh County in 2012. The surrounding area consists of rocky steppe habitat dominated by mountain almond ('' Prunus scoparia''), ''
Astragalus Astragalus may refer to: * ''Astragalus'' (plant), a large genus of herbs and small shrubs *Astragalus (bone) The talus (; Latin for ankle or ankle bone; : tali), talus bone, astragalus (), or ankle bone is one of the group of foot bones known ...
'' and ''Artemisia''. In the Hindu Kush, a Pallas's cat was observed sunbathing at the fringe of a rocky high-elevation plain near Dasht-e Nawar in
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan, is a landlocked country located at the crossroads of Central Asia and South Asia. It is bordered by Pakistan to the Durand Line, east and south, Iran to the Afghanistan–Iran borde ...
's Koh-i-Baba range in April 2007. The Pallas's cat was also photographed multiple times in
Bamyan Province Bamyan, also spelled Bamiyan, Bāmīān or Bāmyān (), is one of the thirty-four provinces of Afghanistan with the city of Bamyan as its center, located in central parts of Afghanistan. The terrain in Bamyan is mountainous or semi-mountainous ...
between 2015 and 2017. In
Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
's Qurumber National Park in
Gilgit-Baltistan Gilgit-Baltistan (; ), formerly known as the Northern Areas, is a region administered by Pakistan as an administrative units of Pakistan, administrative territory and consists of the northern portion of the larger Kashmir region, which has b ...
, an individual was recorded on a
ridge A ridge is a long, narrow, elevated geomorphologic landform, structural feature, or a combination of both separated from the surrounding terrain by steep sides. The sides of a ridge slope away from a narrow top, the crest or ridgecrest, wi ...
in a
juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' ( ) of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere as far south ...
dominated forest at in July 2012. In the Transcaspian Region, its presence was first reported in the Kopet Dag mountains and in the vicinity of the Tedzhen and Murghab Rivers in the late 19th century. In
Turkmenistan Turkmenistan is a landlocked country in Central Asia bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the south and southwest and the Caspian Sea to the west. Ash ...
's Sünt-Hasardag Nature Reserve, a camera trap recorded an individual in 2019. The Pallas's cat is allegedly also present in Köpetdag Nature Reserve. Historical records of the Pallas's cat are known in the Surxondaryo Region and Gissar Range along the border of
Tajikistan Tajikistan, officially the Republic of Tajikistan, is a landlocked country in Central Asia. Dushanbe is the capital city, capital and most populous city. Tajikistan borders Afghanistan to the Afghanistan–Tajikistan border, south, Uzbekistan to ...
and
Uzbekistan , image_flag = Flag of Uzbekistan.svg , image_coat = Emblem of Uzbekistan.svg , symbol_type = Emblem of Uzbekistan, Emblem , national_anthem = "State Anthem of Uzbekistan, State Anthem of the Republ ...
. In
Kyrgyzstan Kyrgyzstan, officially the Kyrgyz Republic, is a landlocked country in Central Asia lying in the Tian Shan and Pamir Mountains, Pamir mountain ranges. Bishkek is the Capital city, capital and List of cities in Kyrgyzstan, largest city. Kyrgyz ...
, it is present at high elevations of Sarychat-Ertash State Nature Reserve and in the foothills of the Alay Range. In 2013, a dead female was found in a valley near Engilchek, Kyrgyzstan. In
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a landlocked country primarily in Central Asia, with a European Kazakhstan, small portion in Eastern Europe. It borders Russia to the Kazakhstan–Russia border, north and west, China to th ...
, it inhabits the highlands and steppes of central and east Kazakhstan Region, the periphery of the Betpak-Dala Desert, the northern Balkhash District and the Tarbagatai Mountains. In the South Siberian Mountains, it inhabits grasslands on the Ukok Plateau and in the Altai, Kuray and Saylyugem Mountains. It is also present in Chagan-Uzun and Argut river basins, Mongun-Taiga, Uvs Lake Basin, Sayano-Shushenski Nature Reserve, Tunkinsky National Park, Lake Gusinoye basin and in the interfluves of the Selenga, Chikoy and Khilok rivers. In the eastern
Sayan Mountains The Sayan Mountains (, ; ) are a mountain range in southern Siberia spanning southeastern Russia (Buryatia, Irkutsk Oblast, Krasnoyarsk Krai, Tuva and Khakassia) and northern Mongolia. Before the rapid expansion of the Tsardom of Russia, the mou ...
, its presence was documented for the first time in 1997. In Transbaikal, it inhabits montane steppes at elevations of , where annual rainfall ranges from . In 2013, an individual was observed on the Vitim Plateau. The Pallas's cat inhabits the semi-desert steppe of Ikh Nartiin Chuluu Nature Reserve in Mongolia. In Khustain Nuruu National Park and Gobi Gurvansaikhan National Park, it prefers rocky and rugged habitats that provides cover and camouflage. On the Tibetan plateau, two Pallas's cats were observed in undulating alpine meadow amidst plateau pika (''Ochotona curzoniae'')
colonies A colony is a territory subject to a form of foreign rule, which rules the territory and its indigenous peoples separated from the foreign rulers, the colonizer, and their '' metropole'' (or "mother country"). This separated rule was often or ...
at in
western China Western China ( zh, s=中国西部, l=, labels=no or zh, s=华西, l=, labels=no) is the west of China. It consists of Southwestern China and Northwestern China. In the definition of the Chinese government, Western China covers six provinces ...
's Qumarlêb County in 2001. One of them swam across an
irrigation Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
channel. In Gêrzê County, an individual was sighted in desert steppe habitat at an elevation of in 2005. In 2011, the Pallas's cat was photographed in an alpine meadow in the core area of Sanjiangyuan National Nature Reserve. In Ruoergai, it was observed at several places in habitat that was frequented by pastoralists and their livestock herds. The presence of the Pallas's cat in the Himalayas was first reported in
Ladakh Ladakh () is a region administered by India as a union territory and constitutes an eastern portion of the larger Kashmir region that has been the subject of a Kashmir#Kashmir dispute, dispute between India and Pakistan since 1947 and India an ...
's
Indus The Indus ( ) is a transboundary river of Asia and a trans- Himalayan river of South and Central Asia. The river rises in mountain springs northeast of Mount Kailash in the Western Tibet region of China, flows northwest through the dis ...
valley in 1991. In Changthang Wildlife Sanctuary, Pallas's cats were sighted close by riverbanks at elevations of in 2013 and 2015. In Gangotri National Park, a Pallas's cat was photographed in rocky alpine scrub at in 2019. In
Sikkim Sikkim ( ; ) is a States and union territories of India, state in northeastern India. It borders the Tibet Autonomous Region of China in the north and northeast, Bhutan in the east, Koshi Province of Nepal in the west, and West Bengal in the ...
, an individual was observed on a rocky slope at an elevation of in the vicinity of Tso Lhamo Lake in 2007. In December 2012, the Pallas's cat was recorded for the first time in the
Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
Himalayas. It was photographed in the upper Marshyangdi river valley in alpine
pasture Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Types of pasture Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, c ...
s at elevations of and in
Annapurna Conservation Area Annapurna Conservation Area is Nepal's largest protected area covering in the Annapurna range of the Himalayas. It ranges in elevation from to the peak of Annapurna I at . The conservation area stretches across Manang District, Nepal, Manang, Mu ...
. In Shey-Phoksundo National Park, Pallas's cat scat was detected at in 2016, the globally highest record to date. In January 2012, it was recorded for the first time in
Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...
, namely in rolling hills dominated by
glacial outwash An outwash plain, also called a sandur (plural: ''sandurs''), sandr or sandar, is a plain formed of glaciofluvial deposits due to meltwater outwash at the terminus of a glacier. As it flows, the glacier grinds the underlying rock surface and ...
and alpine steppe vegetation in Wangchuck Centennial National Park. In autumn 2012, it was also photographed at an elevation of in
Jigme Dorji National Park Jigme Dorji National Park (JDNP), named after the late Jigme Dorji Wangchuck, is the second-largest National Park of Bhutan. History It was established in 1974 and stretches over an area of 4316 km2, thereby spanning all three climate z ...
. In 2019, scat samples of two individuals were found in Sagarmatha National Park, providing the first genetic evidence of the cat's presence in the eastern Himalayas.


Behaviour and ecology

The Pallas's cat is solitary. Of nine Pallas's cat kittens observed in captivity, only the two males
scent mark In ethology, territory is the sociographical area that an animal consistently defends against conspecific competition (or, occasionally, against animals of other species) using agonistic behaviors or (less commonly) real physical aggression. ...
ed by spraying
urine Urine is a liquid by-product of metabolism in humans and many other animals. In placental mammals, urine flows from the Kidney (vertebrates), kidneys through the ureters to the urinary bladder and exits the urethra through the penile meatus (mal ...
. The Pallas's cat uses caves, rock crevices and marmot burrows as shelter. In central Mongolia, 29 Pallas's cats were fitted with radio collars between June 2005 and October 2007. They used 101 dens during this time, including 39 winter dens, 42 summer dens and 20 dens for raising kittens. The summer and winter dens usually had one entrance with a diameter of . They resided in the summer dens for 2–21 days, and in the winter dens for 2–28 days. Summer and maternal dens were close to rocky habitats with little direct sunlight, whereas winter dens were closer to
ravine A ravine is a landform that is narrower than a canyon and is often the product of streambank erosion. Ravines are typically classified as larger in scale than gullies, although smaller than valleys. Ravines may also be called a cleuch, dell, ...
s. The
home range A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. ...
s of 16 females varied from . The home ranges of nine males varied from and overlapped those of one to four females and partly also those of other males. The sizes of their home ranges decreased in winter. In an unprotected area in central Mongolia, Pallas's cats were mainly
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal (active during dawn), vespertine (biology), vespertine/vespertinal (active during dusk), or both. This is distinguished from diurnalit ...
between May and August, but active by day from September to November. Pallas's cats recorded in four study areas in the western Mongolian Altai mountains were also active during the day, but with a lower frequency at sites where livestock was present.


Hunting and diet

The Pallas's cat is a highly specialised predator of small mammals, which it catches by stalking or ambushing near exits of burrows. It also pulls out
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s with its paws from shallow burrows. In the Altai Mountains, remains of long-tailed ground squirrel (''Urocitellus undulatus''), flat-skulled shrew (''Sorex roboratus''), Pallas's pika (''Ochotona pallasi'') and bird feathers were found near breeding burrows of Pallas's cats. In Transbaikal, it preys on Daurian pika (''Ochotona dauurica''), steppe pika (''O. pusilla''), Daurian ground squirrel (''Spermophilus dauricus'') and young of red-billed chough (''Pyrrhocorax pyrrhocorax''). Scat samples of the Pallas's cat collected in the bufferzone of Khustain Nuruu National Park in central Mongolia contained foremost remains of Daurian pika, Mongolian gerbil (''Meriones unguiculatus''), Mongolian silver vole (''Alticola semicanus'') and remains of
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
birds,
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s and
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are amongst what are possibly the most ancient living groups of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grassh ...
s. Brandt's vole (''Lasiopodomys brandtii'') dominated in the diet of Pallas's cats in Mongolia's Sükhbaatar Province after the irruptive growth of this vole population during 2017 to 2020. Scat found in Shey-Phoksundo National Park contained remains of
pika A pika ( , or ) is a small, mountain-dwelling mammal native to Asia and North America. With short limbs, a very round body, an even coat of fur, and no external tail, they resemble their close relative the rabbit, but with short, rounded ears. ...
species and of
woolly hare The woolly hare (''Lepus oiostolus'') is a species of mammal in the family Leporidae. It is found in western and central China, northern India, and Nepal, where its typical habitat is montane grassland. It has a wide range and is present in some ...
(''Lepus oiostolus''). Remains of a cypriniform fish were found in Pallas's cat scat in Gongga Mountain Nature Reserve.


Reproduction and life cycle

The female is sexually mature at the age of about one year. She is in
estrus The estrous cycle (, originally ) is a set of recurring physiological changes induced by reproductive hormones in females of mammalian subclass Theria. Estrous cycles start after sexual maturity in females and are interrupted by anestrous phas ...
for 26 to 42 hours.
Gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...
lasts 66 to 75 days. A captive male Pallas's cat housed under natural lighting conditions showed increased aggressive and territorial behaviour at the onset of the breeding season, lasting from September to December. Its
blood Blood is a body fluid in the circulatory system of humans and other vertebrates that delivers necessary substances such as nutrients and oxygen to the cells, and transports metabolic waste products away from those same cells. Blood is com ...
contained three times more
testosterone Testosterone is the primary male sex hormone and androgen in Male, males. In humans, testosterone plays a key role in the development of Male reproductive system, male reproductive tissues such as testicles and prostate, as well as promoting se ...
than in the non-breeding season, and its
ejaculate Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the penis through the urethra. It is the final stage and natural objective of male sexual stimulation, and an essential component of natural concepti ...
was more concentrated with more normal
sperm Sperm (: sperm or sperms) is the male reproductive Cell (biology), cell, or gamete, in anisogamous forms of sexual reproduction (forms in which there is a larger, female reproductive cell and a smaller, male one). Animals produce motile sperm ...
forms and a higher
motility Motility is the ability of an organism to move independently using metabolism, metabolic energy. This biological concept encompasses movement at various levels, from whole organisms to cells and subcellular components. Motility is observed in ...
of sperm. In the wild, the female gives birth to a
litter Litter consists of waste products that have been discarded incorrectly, without consent, at an unsuitable location. The waste is objects, often man-made, such as aluminum cans, paper cups, food wrappers, cardboard boxes or plastic bottles, but ...
of two to six kittens between the end of April and late May. The newborn kittens' fur is fuzzy, and their eyes are closed until the age of about two weeks. A newborn male kitten born in a zoo weighed , measured and had a long tail. In central Mongolia, seven females with kittens were observed using 20 dens for 4–60 days. Their maternal dens were either among rocks, or in former burrows of the Tarbagan marmot (''Marmota sibirica''), and had at least two entrances. In Iran, a Pallas's cat was observed using cavities of aged Greek juniper ('' Juniperus excelsa'') as breeding dens for a litter of four kittens. Two-month-old kittens weigh , and their fur gradually grows longer. They start hunting at the age of about five months and reach adult size by the age of six to seven months.


Threats

In China, Mongolia and Russia, the Pallas's cat was once hunted for its fur in large numbers of more than 10,000 skins annually. In China and the former
Soviet Union The Union of Soviet Socialist Republics. (USSR), commonly known as the Soviet Union, was a List of former transcontinental countries#Since 1700, transcontinental country that spanned much of Eurasia from 1922 until Dissolution of the Soviet ...
, hunting of the Pallas's cat decreased in the 1970s when it became legally protected. Mongolia exported 9,185 skins in 1987, but international trade has ceased since 1988. However, domestic trade of its skins and body parts for medicinal purposes continues in the country, and it may be hunted throughout the year. Cases of
herding dog A herding dog, also known as a stock dog or working dog, is a Dog type, type of dog that either has been trained in herding livestock or belongs to one of the dog breed, breeds that were developed for herding. A dog specifically trained to he ...
s killing Pallas's cats were reported in Iran, Kazakhstan and the Altai Republic. Pallas's cats have also fallen victim in traps set for small mammals in Kazakhstan and in the Altai Republic. In Transbaikal, the Pallas's cat is threatened by
poaching Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...
. In Mongolia, the use of the
rodenticide Rodenticides are chemicals made and sold for the purpose of killing rodents. While commonly referred to as "rat poison", rodenticides are also used to kill mice, woodchucks, chipmunks, porcupines, nutria, beavers, and voles. Some rodenticides ...
bromadiolone in the frame of rodent control measures in the early 21st century poisoned the prey base of
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they ar ...
s and raptors. In the Sanjiangyuan region of the Tibetan Plateau, of grassland was poisoned between 2005 and 2009, leading to an estimated loss of of pika
biomass Biomass is a term used in several contexts: in the context of ecology it means living organisms, and in the context of bioenergy it means matter from recently living (but now dead) organisms. In the latter context, there are variations in how ...
. The Pallas's cat may be negatively affected by habitat fragmentation due to mining and infrastructure projects.


Conservation

On the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
, the Pallas's cat is classified as Least Concern since 2020 because of its wide-spread range and assumed large global population. It is listed in
CITES Appendix II CITES (shorter acronym for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of inte ...
. Hunting it is prohibited in all range countries except Mongolia. Since 2009, it is legally protected in Afghanistan, where all hunting and trade with its body parts is banned. On the Mongolian Red List of Mammals, it is listed as
Near Threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
since 2006. In China, it is listed as
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
. In Turkmenistan, it is proposed to be listed as
Critically Endangered An IUCN Red List critically endangered (CR or sometimes CE) species is one that has been categorized by the International Union for Conservation of Nature as facing an extremely high risk of extinction in the wild. As of December 2023, of t ...
due to the scarcity of contemporary records.


In captivity

Between 1951 and 1979, the
Beijing Zoo Beijing Zoo is a zoological park in Xizhimen, Xicheng District, Beijing. Founded in 1906 during the late Qing dynasty, it is the oldest zoo in China and oldest public park in northern China. The zoo is also a center of zoological research that ...
kept 16 Pallas's cats, but they lived for less than three years. In 1984, the Pallas's cat was designated as a priority species for
captive breeding Captive breeding, also known as captive propagation, is the process of keeping plants or animals in controlled environments, such as wildlife reserves, zoos, Botanical garden, botanic gardens, and other Conservation biology, conservation facilitie ...
of the American Association of Zoos and Aquariums's Species Survival Plan. Almost half of the kittens born in member zoos died within the first 30 days, reaching the highest
mortality rate Mortality rate, or death rate, is a measure of the number of deaths (in general, or due to a specific cause) in a particular Statistical population, population, scaled to the size of that population, per unit of time. Mortality rate is typically ...
in captivity of any small wild cat. Zoos in the former Soviet Union received most of the wild-caught Pallas's cats from the Transbaikal region and a few from Mongolia.
Moscow Zoo The Moscow Zoo or Moskovsky Zoopark () is a zoo, the largest in Russia. History The Moscow Zoo was founded in 1864 by professor-biologists, K.F. Rulje, S.A. Usov and A.P. Bogdanov, from the Moscow State University. In 1919, the zoo was natio ...
initiated a
studbook A breed registry, also known as a herdbook, studbook or register, in animal husbandry, the hobby of animal fancy, is an official list of animals within a specific breed whose parents are known. Animals are usually registered by their breeders w ...
for the Pallas's cat in 1997. Since 2004, the Pallas's cat international studbook has been managed by the Royal Zoological Society of Scotland, which also coordinates the captive breeding program for the Pallas's cat within the
European Endangered Species Programme The EAZA Ex-situ Programme (EEP) is a population management and Ex situ conservation, conservation programme by European Association of Zoos and Aquaria, European Association of Zoos and Aquaria (EAZA) for wild animals living in European zoos. The p ...
. As of 2018, 177 Pallas's cats were kept in 60 zoos in Europe, Russia, North America and Japan. In 2011, a female Pallas's cat was artificially inseminated for the first time with
semen Semen, also known as seminal fluid, is a bodily fluid that contains spermatozoon, spermatozoa which is secreted by the male gonads (sexual glands) and other sexual organs of male or hermaphrodite, hermaphroditic animals. In humans and placen ...
from the male at the Cincinnati Zoo. After 69 days, she gave birth to four kittens, of which one was stillborn.


In popular culture

The Pallas's cat is featured in a Russian
Internet meme An Internet meme, or meme (, Help:Pronunciation respelling key, ''MEEM''), is a cultural item (such as an idea, behavior, or style) that spreads across the Internet, primarily through Social media, social media platforms. Internet memes manif ...
known as "Pet the cat" introduced in 2008; the meme is a picture of a Pallas's cat that invites the reader to pet it in the image's caption. In 2012, the Pallas's cat became the mascot of Moscow Zoo.


References


External links

* * * * {{DEFAULTSORT:cat, Pallas's Pallas's cat Pallas's cat IUCN Red List least concern species Pallas's cat Taxa named by Peter Simon Pallas