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botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
, a fruit is the
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
-bearing structure in
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s (angiosperms) that is formed from the
ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
after
flowering Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propagated using the movements of humans and other animals in a
symbiotic relationship Symbiosis (Ancient Greek : living with, companionship < : together; and ''bíōsis'': living) is any type of a close and long-term biolo ...
that is the means for
seed dispersal In spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
for the one group and
nutrition Nutrition is the biochemistry, biochemical and physiology, physiological process by which an organism uses food and water to support its life. The intake of these substances provides organisms with nutrients (divided into Macronutrient, macro- ...
for the other; humans, and many other animals, have become dependent on fruits as a source of food. Consequently, fruits account for a substantial fraction of the world's
agricultural Agriculture encompasses crop and livestock production, aquaculture, and forestry for food and non-food products. Agriculture was a key factor in the rise of sedentary human civilization, whereby farming of domesticated species created f ...
output, and some (such as the
apple An apple is a round, edible fruit produced by an apple tree (''Malus'' spp.). Fruit trees of the orchard or domestic apple (''Malus domestica''), the most widely grown in the genus, are agriculture, cultivated worldwide. The tree originated ...
and the
pomegranate The pomegranate (''Punica granatum'') is a fruit-bearing deciduous shrub in the family Lythraceae, subfamily Punica, Punicoideae, that grows between tall. Rich in symbolic and mythological associations in many cultures, it is thought to have o ...
) have acquired extensive cultural and symbolic meanings. In common language and culinary usage, ''fruit'' normally means the seed-associated fleshy structures (or produce) of plants that typically are sweet (or sour) and edible in the raw state, such as apples, bananas, grapes, lemons, oranges, and strawberries. In botanical usage, the term ''fruit'' also includes many structures that are not commonly called as such in everyday language, such as nuts, bean pods, corn kernels, tomatoes, and wheat grains.


Botanical vs. culinary

Many common language terms used for fruit and seeds differ from botanical classifications. For example, in botany, a ''fruit'' is a ripened
ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
or
carpel Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more ...
that contains seeds, e.g., an orange, pomegranate, tomato or a pumpkin. A '' nut'' is a type of fruit (and not a seed), and a ''seed'' is a ripened
ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the sporangium, megasporangium), ...
. In culinary language, a ''fruit'' is the sweet- or not sweet- (even sour-) tasting produce of a specific plant (e.g., a peach, pear or lemon); ''nuts'' are hard, oily, non-sweet plant produce in shells (
hazelnut The hazelnut is the fruit of the hazel tree and therefore includes any of the nuts deriving from species of the genus '' Corylus'', especially the nuts of the species ''Corylus avellana''. They are also known as cobnuts or filberts according to ...
,
acorn The acorn is the nut (fruit), nut of the oaks and their close relatives (genera ''Quercus'', ''Notholithocarpus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains a seedling surrounded by two cotyledons (seedling leaves), en ...
). ''
Vegetables Vegetables are edible parts of plants that are consumed by humans or other animals as food. This original meaning is still commonly used, and is applied to plants collectively to refer to all edible plant matter, including flowers, fruits, ...
'', so-called, typically are savory or non-sweet produce (
zucchini Zucchini (; : ''zucchini'' or ''zucchinis''), courgette () or ''Cucurbita pepo'' is a summer squash, a Vine, vining herbaceous plant whose fruit are harvested when their immature seeds and Fruit anatomy#Epicarp, epicarp (rind) are still soft a ...
, lettuce, broccoli, and tomato). but some may be sweet-tasting (sweet potato). Examples of botanically classified fruit that are typically called vegetables include
cucumber The cucumber (''Cucumis sativus'') is a widely-cultivated creeping vine plant in the family Cucurbitaceae that bears cylindrical to spherical fruits, which are used as culinary vegetables.pumpkin A pumpkin is a cultivar, cultivated winter squash in the genus ''Cucurbita''. The term is most commonly applied to round, orange-colored squash varieties, but does not possess a scientific definition. It may be used in reference to many dif ...
, and squash (all are
cucurbits The Cucurbitaceae (), also called cucurbits or the gourd family, are a plant family consisting of about 965 species in 101 genera.
);
bean A bean is the seed of some plants in the legume family (Fabaceae) used as a vegetable for human consumption or animal feed. The seeds are often preserved through drying (a ''pulse''), but fresh beans are also sold. Dried beans are traditi ...
s,
peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), goober pea, pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible seeds. It is widely grown in the tropics and subtropics by small and large ...
s, and
pea Pea (''pisum'' in Latin) is a pulse or fodder crop, but the word often refers to the seed or sometimes the pod of this flowering plant species. Peas are eaten as a vegetable. Carl Linnaeus gave the species the scientific name ''Pisum sativum' ...
s (all
legumes Legumes are plants in the pea family Fabaceae (or Leguminosae), or the fruit or seeds of such plants. When used as a dry grain for human consumption, the seeds are also called pulses. Legumes are grown agriculturally, primarily for human consu ...
); and
corn Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout Poaceae, grass that produces cereal grain. It was domesticated by indigenous peoples of Mexico, indigenous peoples in southern Mexico about 9,000 years ago ...
,
eggplant Eggplant (American English, US, Canadian English, CA, Australian English, AU, Philippine English, PH), aubergine (British English, UK, Hiberno English, IE, New Zealand English, NZ), brinjal (Indian English, IN, Singapore English, SG, Malays ...
,
bell pepper The bell pepper (also known as sweet pepper, paprika, pepper, capsicum or, in some parts of the US midwest, mango) is the fruit of plants in the Grossum Group of the species ''Capsicum annuum''. Cultivars of the plant produce fruits in diff ...
(or sweet pepper), and
tomato The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible Berry (botany), berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originate ...
. Many spices are fruits, botanically speaking, including
black pepper Black pepper (''Piper nigrum'') is a flowering vine in the family Piperaceae, cultivated for its fruit (the peppercorn), which is usually dried and used as a spice and seasoning. The fruit is a drupe (stonefruit) which is about in diameter ...
,
chili pepper Chili peppers, also spelled chile or chilli ( ), are varieties of fruit#Berries, berry-fruit plants from the genus ''Capsicum'', which are members of the nightshade family Solanaceae, cultivated for their pungency. They are used as a spice to ...
,
cumin Cumin (, ; ; ''Cuminum cyminum'') is a flowering plant in the family Apiaceae, native to the Irano-Turanian Region. Its seeds – each one contained within a fruit, which is dried – are used in the cuisines of many cultures in both whole ...
and
allspice Allspice, also known as Jamaica pepper, myrtle pepper, pimenta, or pimento, is the dried unripe berry of ''Pimenta dioica'', a midcanopy tree native to the Greater Antilles, southern Mexico, and Central America, now cultivated in many warm par ...
. In contrast,
rhubarb Rhubarb is the fleshy, edible stalks ( petioles) of species and hybrids (culinary rhubarb) of ''Rheum'' in the family Polygonaceae, which are cooked and used for food. The plant is a herbaceous perennial that grows from short, thick rhizomes. ...
is often called a fruit when used in making
pies A pie is a Baking, baked dish which is usually made of a pastry dough casing that contains a filling of various sweetness, sweet or Savoury taste, savoury ingredients. Sweet pies may be filled with fruit (as in an apple pie), nuts (pecan pie) ...
, but the edible produce of rhubarb is actually the leaf stalk or petiole of the plant. Edible
gymnosperm The gymnosperms ( ; ) are a group of woody, perennial Seed plant, seed-producing plants, typically lacking the protective outer covering which surrounds the seeds in flowering plants, that include Pinophyta, conifers, cycads, Ginkgo, and gnetoph ...
seeds are often given fruit names, e.g.,
ginkgo ''Ginkgo'' is a genus of non-flowering seed plants, assigned to the gymnosperms. The scientific name is also used as the English common name. The order to which the genus belongs, Ginkgoales, first appeared in the Permian, , and ''Ginkgo'' is n ...
nuts and
pine nut Pine nuts, also called piñón (), pinoli (), or pignoli, are the edible seeds of pines (family Pinaceae, genus ''Pinus''). According to the Food and Agriculture Organization, only 29 species provide edible nuts, while 20 are traded locall ...
s. Botanically, a
cereal A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...
grain, such as
corn Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout Poaceae, grass that produces cereal grain. It was domesticated by indigenous peoples of Mexico, indigenous peoples in southern Mexico about 9,000 years ago ...
,
rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...
, or
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
is a kind of fruit (termed a
caryopsis In botany, a caryopsis () is a type of simple fruit—one that is monocarpellate (formed from a single carpel) and indehiscent (not opening at maturity) and resembles an achene, except that in a caryopsis the pericarp is fused with the thin s ...
). However, the fruit wall is thin and fused to the seed coat, so almost all the edible grain-fruit is actually a seed.


Structure

The outer layer, often edible, of most fruits is called the ''pericarp''. Typically formed from the ovary, it surrounds the seeds; in some species, however, other structural tissues contribute to or form the edible portion. The pericarp may be described in three layers from outer to inner, i.e., the ''epicarp'', ''mesocarp'' and ''endocarp''. Fruit that bears a prominent pointed terminal projection is said to be ''beaked''.


Development

A fruit results from the fertilizing and maturing of one or more flowers. The
gynoecium Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
, which contains the '' stigma-style-ovary'' system, is centered in the flower-head, and it forms all or part of the fruit. Inside the ovary(ies) are one or more
ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the sporangium, megasporangium), ...
s. Here begins a complex sequence called ''
double fertilization Double fertilization or double fertilisation (see spelling differences) is a complex fertilization mechanism of angiosperms. This process involves the fusion of a female gametophyte or megagametophyte, also called the embryonic sac, with two m ...
'': a female
gametophyte A gametophyte () is one of the two alternating multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the se ...
produces an egg cell for the purpose of fertilization. (A female gametophyte is called a '' megagametophyte'', and also called the ''
embryo sac In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...
''.) After double fertilization, the ovules will become seeds. Ovules are fertilized in a process that starts with
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or bu ...
, which is the movement of pollen from the stamens to the stigma-style-ovary system within the flower-head. After pollination, a
pollen tube A pollen tube is a tubular structure produced by the male gametophyte of seed plants when it germinates. Pollen tube elongation is an integral stage in the plant life cycle. The pollen tube acts as a conduit to transport the male gamete cells fr ...
grows from the (deposited) pollen through the stigma down the style into the ovary to the ovule. Two sperm are transferred from the pollen to a megagametophyte. Within the megagametophyte, one sperm unites with the egg, forming a
zygote A zygote (; , ) is a eukaryote, eukaryotic cell (biology), cell formed by a fertilization event between two gametes. The zygote's genome is a combination of the DNA in each gamete, and contains all of the genetic information of a new individ ...
, while the second sperm enters the central cell forming the endosperm mother cell, which completes the double fertilization process. Later, the zygote will give rise to the embryo of the seed, and the endosperm mother cell will give rise to
endosperm The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the Embryo#Pla ...
, a nutritive tissue used by the embryo. Fruit formation is associated with
meiosis Meiosis () is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, the sperm or egg cells. It involves two rounds of division that ultimately result in four cells, each with only one c ...
, a central aspect of sexual reproduction in flowering plants. During meiosis
homologous chromosome Homologous chromosomes or homologs are a set of one maternal and one paternal chromosome that pair up with each other inside a cell during meiosis. Homologs have the same genes in the same locus (genetics), loci, where they provide points along e ...
s replicate, recombine and randomly segregate, and then undergo segregation of
sister chromatids A sister chromatid refers to the identical copies ( chromatids) formed by the DNA replication of a chromosome, with both copies joined together by a common centromere. In other words, a sister chromatid may also be said to be 'one-half' of the du ...
to produce
haploid Ploidy () is the number of complete sets of chromosomes in a cell (biology), cell, and hence the number of possible alleles for Autosome, autosomal and Pseudoautosomal region, pseudoautosomal genes. Here ''sets of chromosomes'' refers to the num ...
cells. Union of haploid nuclei from
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
and
ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the sporangium, megasporangium), ...
(
fertilisation Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a zygote and initiate its development into a new individual organism or of ...
), occurring either by
self In philosophy, the self is an individual's own being, knowledge, and values, and the relationship between these attributes. The first-person perspective distinguishes selfhood from personal identity. Whereas "identity" is (literally) same ...
- or cross-
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or bu ...
, leads to the formation of a diploid
zygote A zygote (; , ) is a eukaryote, eukaryotic cell (biology), cell formed by a fertilization event between two gametes. The zygote's genome is a combination of the DNA in each gamete, and contains all of the genetic information of a new individ ...
that can then develop into an
embryo An embryo ( ) is the initial stage of development for a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sp ...
within the emerging seed. Repeated fertilisations within the
ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
are accompanied by maturation of the ovary to form the fruit. As the ovules develop into seeds, the ovary begins to ripen and the ovary wall, the ''pericarp'', may become fleshy (as in berries or
drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
s), or it may form a hard outer covering (as in nuts). In some multi-seeded fruits, the extent to which a fleshy structure develops is proportional to the number of fertilized ovules. The pericarp typically is differentiated into two or three distinct layers; these are called the ''exocarp'' (outer layer, also called epicarp), ''mesocarp'' (middle layer), and ''endocarp'' (inner layer). In some fruits, the
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s,
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s,
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s or the style of the flower fall away as the fleshy fruit ripens. However, for simple fruits derived from an ''
inferior ovary In the flowering plants, an ovary is a part of the female reproductive organ of the flower or gynoecium. Specifically, it is the part of the pistil which holds the ovule(s) and is located above or below or at the point of connection with the ba ...
'' – i.e., one that lies the attachment of other floral parts – there are parts (including petals, sepals, and stamens) that fuse with the ovary and ripen with it. For such a case, when floral parts other than the ovary form a significant part of the fruit that develops, it is called an
accessory fruit An accessory fruit is a fruit that contains tissue derived from plant parts other than the Ovary (botany), ovary. In other words, the flesh of the fruit develops not from the floral ovary, but from some adjacent tissue exterior to the carpel (f ...
. Examples of accessory fruits include apple, rose hip, strawberry, and pineapple. Because several parts of the flower besides the ovary may contribute to the structure of a fruit, it is important to understand how a particular fruit forms. There are three general modes of fruit development: * Apocarpous fruits develop from a ''single flower'' (while having one or more separate, unfused, carpels); they are the simple fruits. *
Syncarpous Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ...
fruits develop from a ''single
gynoecium Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
'' (having two or more carpels fused together). * Multiple fruits form from many flowers – i.e., an inflorescence of flowers. File:Nectarine Fruit Development.jpg, The development sequence of a typical
drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
, the
nectarine The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in China. It bears edible juicy fruits with various characteristics, most called peaches and the glossy-skinned, non-fuzzy varieties called nectarines. Peach ...
(''Prunus persica'') over a 7.5-month period, from bud formation in early winter to fruit
ripening Ripening is a process in fruits that causes them to become more palatable. In general, fruit becomes sweeter, less green, and softer as it ripens. Even though the acidity of fruit increases as it ripens, the higher acidity level does not make t ...
in midsummer File:Mature flower diagram.svg, The parts of a flower, showing the stigma-style-ovary system. File:Pome apples text.jpg, An apple is a simple, fleshy fruit. Key parts are the epicarp, or exocarp, or outer skin (not labelled); and the mesocarp and endocarp (labelled). image:Ovary position.svg, upright 1.5, Insertion point: There are three positions of insertion of the ovary at the base of a flower: I superior; II half-inferior; III inferior. The 'insertion point' is where the
androecium The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
parts (a), the petals (p), and the sepals (s) all converge and attach to the receptacle (r). (Ovary=gynoecium (g).) File:Noni fruit dev.jpg, upright 1.5, In the
noni ''Morinda citrifolia'' is a fruit-bearing tree in the coffee family, Rubiaceae, native to Southeast Asia and Australasia, which was spread across the Pacific by Polynesian sailors. The species is now cultivated throughout the tropics and widel ...
, flowers are produced in time-sequence along the stem. It is possible to see a progression of flowering, fruit development, and fruit ripening. Tweelinggroei appels.jpg, Twin apples


Classification of fruits

Consistent with the three modes of fruit development, plant scientists have classified fruits into three main groups: simple fruits, aggregate fruits, and multiple (or composite) fruits. The groupings reflect how the ovary and other flower organs are arranged and how the fruits develop, but they are not evolutionarily relevant as diverse plant
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
may be in the same group. While the section of a
fungus A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
that produces
spore In biology, a spore is a unit of sexual reproduction, sexual (in fungi) or asexual reproduction that may be adapted for biological dispersal, dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores fo ...
s is called a ''fruiting'' body, fungi are members of the fungi kingdom and not of the plant kingdom.


Simple fruits

Simple fruits are the result of the ripening-to-fruit of a simple or compound ovary in a ''single flower'' with a ''single
pistil Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
''. In contrast, a single flower with numerous pistils typically produces an
aggregate fruit A raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupelets. image:Aquilegia vulgaris 004.JPG, The fruit of an ''Aquilegi ...
; and the merging of several flowers, or a 'multiple' of flowers, results in a 'multiple' fruit. A simple fruit is further classified as either dry or fleshy. To distribute their seeds, dry fruits may split open and discharge their seeds to the winds, which is called dehiscence. Or the distribution process may rely upon the decay and degradation of the fruit to expose the seeds; or it may rely upon the eating of fruit and excreting of seeds by
frugivores A frugivore ( ) is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance ...
– both are called indehiscence. Fleshy fruits do not split open, but they also are indehiscent and they may also rely on frugivores for distribution of their seeds. Typically, the entire outer layer of the ovary wall ripens into a potentially edible
pericarp Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Fruitlike structures may develop directly from the seed itself rather th ...
. Types of dry simple fruits, (with examples) include: *
Achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple fruit, simple dry fruits, dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and Dehi ...
– most commonly seen in aggregate fruits (e.g.,
strawberry The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown Hybrid (biology), hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit ...
, see below). * Capsule – (
Brazil nut The Brazil nut (''Bertholletia excelsa'') is a South American tree in the family Lecythidaceae, and it is also the name of the tree's commercially harvested edible seeds. It is one of the largest and longest-lived trees in the Amazon rainforest. ...
: botanically, it is not a nut). *
Caryopsis In botany, a caryopsis () is a type of simple fruit—one that is monocarpellate (formed from a single carpel) and indehiscent (not opening at maturity) and resembles an achene, except that in a caryopsis the pericarp is fused with the thin s ...
– (cereal grains, including
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
,
rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...
,
oats The oat (''Avena sativa''), sometimes called the common oat, is a species of cereal grain grown for its seed, which is known by the same name (usually in the plural). Oats appear to have been domesticated as a secondary crop, as their seed ...
,
barley Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...
). * Cypsela – an achene-like fruit derived from the individual florets in a
capitulum capitulum (plural capitula) may refer to: *the Latin word for chapter ** an index or list of chapters at the head of a gospel manuscript ** a short reading in the Liturgy of the Hours *** derived from which, it is the Latin for the assembly known ...
: (
dandelion ''Taraxacum'' () is a genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus has a near-cosmopolitan distribu ...
). * Fibrous drupe – (
coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family (biology), family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, ...
,
walnut A walnut is the edible seed of any tree of the genus '' Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. They are accessory fruit because the outer covering of the fruit is technically an i ...
: botanically, neither is a true nut.). * Follicle – follicles are formed from a single carpel, and opens by one suture: (
milkweed ''Asclepias'' is a genus of herbaceous, perennial, flowering plants known as milkweeds, named for their latex, a milky substance containing cardiac glycosides termed cardenolides, exuded where cells are damaged. Most species are toxic to huma ...
); also commonly seen in aggregate fruits: (
magnolia ''Magnolia'' is a large genus of about 210 to 340The number of species in the genus ''Magnolia'' depends on the taxonomic view that one takes up. Recent molecular and morphological research shows that former genera ''Talauma'', ''Dugandiodendr ...
,
peony The peony or paeony () is any flowering plant in the genus ''Paeonia'', the only genus in the family Paeoniaceae. Peonies are native to Asia, Europe, and Western North America. Scientists differ on the number of species that can be distinguish ...
). *
Legume Legumes are plants in the pea family Fabaceae (or Leguminosae), or the fruit or seeds of such plants. When used as a dry grain for human consumption, the seeds are also called pulses. Legumes are grown agriculturally, primarily for human consum ...
– (
bean A bean is the seed of some plants in the legume family (Fabaceae) used as a vegetable for human consumption or animal feed. The seeds are often preserved through drying (a ''pulse''), but fresh beans are also sold. Dried beans are traditi ...
,
pea Pea (''pisum'' in Latin) is a pulse or fodder crop, but the word often refers to the seed or sometimes the pod of this flowering plant species. Peas are eaten as a vegetable. Carl Linnaeus gave the species the scientific name ''Pisum sativum' ...
,
peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), goober pea, pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible seeds. It is widely grown in the tropics and subtropics by small and large ...
: botanically, the peanut is the seed of a legume, not a nut). *
Loment A loment (or lomentum) is a part of certain legume plants. It is a type of dehiscent fruit that breaks apart at the constrictions occurring between segments, so that each segment contains only one seed. It is a type of schizocarp. Tick trefoil ('' ...
– a type of
indehiscent Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part. Structures that ...
legume: ( sweet vetch or wild potato). * Nut – ( beechnut,
hazelnut The hazelnut is the fruit of the hazel tree and therefore includes any of the nuts deriving from species of the genus '' Corylus'', especially the nuts of the species ''Corylus avellana''. They are also known as cobnuts or filberts according to ...
,
acorn The acorn is the nut (fruit), nut of the oaks and their close relatives (genera ''Quercus'', ''Notholithocarpus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains a seedling surrounded by two cotyledons (seedling leaves), en ...
(of the
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
): botanically, these are true nuts). *
Samara Samara, formerly known as Kuybyshev (1935–1991), is the largest city and administrative centre of Samara Oblast in Russia. The city is located at the confluence of the Volga and the Samara (Volga), Samara rivers, with a population of over 1.14 ...
– ( ash,
elm Elms are deciduous and semi-deciduous trees comprising the genus ''Ulmus'' in the family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical- montane regions of North America and Eurasia, ...
, maple key). *
Schizocarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any Dry fruits, dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more ...
, see below – ( carrot seed). *
Silique A silique or siliqua (plural ''siliques'' or ''siliquae'') is a type of fruit (seed capsule) having two fused carpels with the length being more than three times the width. When the length is less than three times the width of the dried fruit ...
– (
radish The radish (''Raphanus sativus'') is a flowering plant in the mustard family, Brassicaceae. Its large taproot is commonly used as a root vegetable, although the entire plant is edible and its leaves are sometimes used as a leaf vegetable. Origina ...
seed). * Silicle – (
shepherd's purse ''Capsella bursa-pastoris'', known as shepherd's purse or lady's purse because of its triangular flat fruits, which are purse-like, is a small annual and ruderal flowering plant in the mustard family (Brassicaceae). Scientists have referred to ...
). * Utricle – (
beet The beetroot (British English) or beet (North American English) is the taproot portion of a '' Beta vulgaris'' subsp. ''vulgaris'' plant in the Conditiva Group. The plant is a root vegetable also known as the table beet, garden beet, dinner ...
, ''
Rumex The docks and sorrels, genus ''Rumex'', are a genus of about 200 species of annual, biennial, and perennial herbs in the buckwheat family, Polygonaceae. Members of this genus are very common perennial herbs with a native almost worldwide distri ...
''). Fruits in which part or all of the ''pericarp'' (fruit wall) is fleshy at maturity are termed ''fleshy simple fruits''. Types of fleshy simple fruits, (with examples) include: *
Berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...
– the berry is the most common type of fleshy fruit. The entire outer layer of the ovary wall ripens into a potentially edible "pericarp", (see below). *
Stone fruit In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
or drupe – the definitive characteristic of a drupe is the hard, "lignified" stone (sometimes called the "pit"). It is derived from the ovary wall of the flower:
apricot An apricot (, ) is a fruit, or the tree that bears the fruit, of several species in the genus ''Prunus''. Usually an apricot is from the species '' P. armeniaca'', but the fruits of the other species in ''Prunus'' sect. ''Armeniaca'' are also ...
,
cherry A cherry is the fruit of many plants of the genus ''Prunus'', and is a fleshy drupe (stone fruit). Commercial cherries are obtained from cultivars of several species, such as the sweet '' Prunus avium'' and the sour '' Prunus cerasus''. The na ...
,
olive The olive, botanical name ''Olea europaea'' ("European olive"), is a species of Subtropics, subtropical evergreen tree in the Family (biology), family Oleaceae. Originating in Anatolia, Asia Minor, it is abundant throughout the Mediterranean ...
,
peach The peach (''Prunus persica'') is a deciduous tree first domesticated and Agriculture, cultivated in China. It bears edible juicy fruits with various characteristics, most called peaches and the glossy-skinned, non-fuzzy varieties called necta ...
,
plum A plum is a fruit of some species in Prunus subg. Prunus, ''Prunus'' subg. ''Prunus'.'' Dried plums are often called prunes, though in the United States they may be labeled as 'dried plums', especially during the 21st century. Plums are ...
,
mango A mango is an edible stone fruit produced by the tropical tree '' Mangifera indica''. It originated from the region between northwestern Myanmar, Bangladesh, and northeastern India. ''M. indica'' has been cultivated in South and Southeast As ...
. *
Pome In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Pome fruits consist of a central "core" containing multiple small seeds, which is enveloped by a tough membrane and surrounded by a ...
– the pome fruits:
apple An apple is a round, edible fruit produced by an apple tree (''Malus'' spp.). Fruit trees of the orchard or domestic apple (''Malus domestica''), the most widely grown in the genus, are agriculture, cultivated worldwide. The tree originated ...
s,
pear Pears are fruits produced and consumed around the world, growing on a tree and harvested in late summer into mid-autumn. The pear tree and shrub are a species of genus ''Pyrus'' , in the Family (biology), family Rosaceae, bearing the Pome, po ...
s, rosehips, saskatoon berry, etc., are a
syncarpous Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ...
(fused) fleshy fruit, a simple fruit, developing from a half-inferior ovary. Pomes are of the family
Rosaceae Rosaceae (), the rose family, is a family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus '' Rosa''. The family includes herbs, shrubs, and trees. Most species are deciduous, but som ...
.


Berries

Berries are a type of simple fleshy fruit that issue from a single ovary. (The ovary itself may be compound, with several carpels.) The botanical term ''true berry'' includes grapes, currants, cucumbers, eggplants (aubergines), tomatoes, chili peppers, and bananas, but excludes certain fruits that are called "-berry" by culinary custom or by common usage of the term – such as strawberries and raspberries. Berries may be formed from one or more carpels (i.e., from the simple or compound ovary) from the same, single flower. Seeds typically are embedded in the fleshy interior of the ovary. Examples include: *
Tomato The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible Berry (botany), berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originate ...
– in culinary terms, the tomato is regarded as a vegetable, but it is botanically classified as a fruit and a berry. *
Banana A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large treelike herbaceous flowering plants in the genus '' Musa''. In some countries, cooking bananas are called plantains, distinguishing the ...
– the fruit has been described as a "leathery berry". In cultivated varieties, the seeds are diminished nearly to non-existence. * Pepo – berries with skin that is hardened:
cucurbits The Cucurbitaceae (), also called cucurbits or the gourd family, are a plant family consisting of about 965 species in 101 genera.
, including gourds, squash, melons. *
Hesperidium A hesperidium (: hesperidia) is a modified berry with a tough, leathery rind, as in the oranges and lemons of the genus ''Citrus''. Etymology Carl Linnaeus gave the name Hesperideæ to an order containing the genus ''Citrus'', in allusion to ...
– berries with a rind and a juicy interior: most
citrus ''Citrus'' is a genus of flowering trees and shrubs in the family Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, mandarins, lemons, grapefruits, pomelos, and limes. ''Citrus'' is nativ ...
fruit. *
Cranberry Cranberries are a group of evergreen dwarf shrubs or trailing vines in the subgenus ''Oxycoccus'' of the genus ''Vaccinium''. Cranberries are low, creeping shrubs or vines up to long and in height; they have slender stems that are not th ...
,
gooseberry Gooseberry ( or (American and northern British) or (southern British)) is a common name for many species of ''Ribes'' (which also includes Ribes, currants), as well as a large number of plants of similar appearance, and also several unrela ...
,
redcurrant The redcurrant or red currant (''Ribes rubrum'') is a member of the genus ''Ribes'' in the gooseberry family. It is native to western Europe. The species is widely cultivated and has escaped into the wild in many regions. Description ''Ribes ...
,
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began approximately 8,0 ...
. The strawberry, regardless of its appearance, is classified as a dry, not a fleshy fruit. Botanically, it is not a berry (botany)#Accessory fruits, berry; it is an accessory fruit, aggregate-accessory fruit, the latter term meaning the fleshy part is derived not from the plant's ovaries but from the Receptacle (botany), receptacle that holds the Ovary (botany), ovaries.Esau, K. (1977). ''Anatomy of seed plants''. John Wiley and Sons, New York. Numerous dry achenes are attached to the outside of the fruit-flesh; they appear to be seeds but each is actually an ovary of a flower, with a seed inside.
Schizocarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any Dry fruits, dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more ...
s are dry fruits, though some appear to be fleshy. They originate from syncarpous ovaries but do not actually dehiscence (botany), dehisce; rather, they split into segments with one or more seeds. They include a number of different forms from a wide range of families, including carrot, parsnip, parsley,
cumin Cumin (, ; ; ''Cuminum cyminum'') is a flowering plant in the family Apiaceae, native to the Irano-Turanian Region. Its seeds – each one contained within a fruit, which is dried – are used in the cuisines of many cultures in both whole ...
.


Aggregate fruits

An aggregate fruit is also called an aggregation, or ''etaerio''; it develops from a single flower that presents numerous simple pistils. Each pistil contains one
carpel Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more ...
; together, they form a fruitlet. The ultimate (fruiting) development of the aggregation of pistils is called an ''aggregate fruit'', ''etaerio fruit'', or simply an ''etaerio''. Different types of aggregate fruits can produce different etaerios, such as achenes, drupelets, follicles, and berries. * For example, the Ranunculaceae species, including ''Clematis'' and ''Ranunculus'', produces an etaerio of achenes; * ''Rubus'' species, including raspberry: an etaerio of drupelets; * ''Calotropis'' species: an etaerio of Follicle (fruit), follicles fruit; * ''Annona'' species: an etaerio of berries. Some other broadly recognized species and their etaerios (or aggregations) are: * Teasel; fruit is an aggregation of cypsela (botany), cypselas. * tuliptree#Description, Tuliptree; fruit is an aggregation of samara (fruit), samaras. * Magnolia and peony#Morphology, peony; fruit is an aggregation of follicles. * liquidambar#Species, American sweet gum; fruit is an aggregation of capsules. * Platanus occidentalis#Description, Sycamore; fruit is an aggregation of achenes. The pistils of the raspberry are called ''drupelets'' because each pistil is like a small
drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
attached to the receptacle. In some bramble fruits, such as blackberry, the receptacle, an accessory part, elongates and then develops as part of the fruit, making the blackberry an #Accessory fruit, aggregate-accessory fruit. The strawberry is also an aggregate-accessory fruit, of which the seeds are contained in the achenes. Notably in all these examples, the fruit develops from a single flower, with numerous pistils.


Hybrid fruits

Hybrid fruits are created through the controlled speciation of fruits that creates new varieties and cross-breeds. Hybrid (biology), Hybrids are grown using plant propagation to create new cultivars. This may introduce an entirely new type of fruit or improve the properties of an existing fruit.


Examples

* Nectaplum * Pluot * Tangelo * Orangequat * Strawberry, Garden strawberry * Plumcot * Rangpur (fruit), Rangpur * Blood lime * Ugli fruit * Tayberry


Multiple fruits

A multiple fruit is formed from a cluster of flowers, (a 'multiple' of flowers) – also called an ''inflorescence''. Each ('smallish') flower produces a single fruitlet, which, as all develop, all merge into one mass of fruit. Examples include pineapple, ficus, fig, mulberry, Osage orange, and breadfruit. An inflorescence (a cluster) of white flowers, called a head, is produced first. After Fertilization#Fertilisation in plants, fertilization, each flower in the cluster develops into a drupe; as the drupes expand, they develop as a ''connation, connate'' organ, merging into a multiple fleshy fruit called a ''syncarp''. Progressive stages of multiple flowering and fruit development can be observed on a single branch of the Indian mulberry, or ''
noni ''Morinda citrifolia'' is a fruit-bearing tree in the coffee family, Rubiaceae, native to Southeast Asia and Australasia, which was spread across the Pacific by Polynesian sailors. The species is now cultivated throughout the tropics and widel ...
''. During the sequence of development, a progression of second, third, and more inflorescences are initiated in turn at the head of the branch or stem.


Accessory fruit forms

Fruits may incorporate tissues derived from other floral parts besides the ovary, including the receptacle, hypanthium, petals, or sepals. Accessory fruits occur in all three classes of fruit development – simple, aggregate, and multiple. Accessory fruits are frequently designated by the hyphenated term showing both characters. For example, a pineapple is a multiple-accessory fruit, a blackberry is an aggregate-accessory fruit, and an apple is a simple-accessory fruit.


Table of fleshy fruit examples


Seedless fruits

Seedlessness is an important feature of some fruits of commerce. Commercial cultivars of bananas and pineapples are examples of seedless fruits. Some cultivars of
citrus ''Citrus'' is a genus of flowering trees and shrubs in the family Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, mandarins, lemons, grapefruits, pomelos, and limes. ''Citrus'' is nativ ...
fruits (especially grapefruit, mandarin oranges, navel oranges, Mikan, satsumas), table grapes, and of watermelons are valued for their seedlessness. In some species, seedlessness is the result of ''parthenocarpy'', where fruits set without fertilization. Parthenocarpic fruit-set may (or may not) require pollination, but most seedless citrus fruits require a stimulus from pollination to produce fruit. Seedless bananas and grapes are triploids, and seedlessness results from the abortion of the
embryo An embryo ( ) is the initial stage of development for a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sp ...
nic plant that is produced by fertilization, a phenomenon known as ''stenospermocarpy'', which requires normal pollination and fertilization.


Seed dispersal

Variations in fruit structures largely depend on the Biological dispersal, modes of dispersal applied to their seeds. Dispersal is achieved by wind or water, by explosive dehiscence, and by interactions with animals. Some fruits present their outer skins or shells coated with spikes or hooked burrs; these evolved either to deter would-be foragers from feeding on them or to serve to attach themselves to the hair, feathers, legs, or clothing of animals, thereby using them as dispersal agents. These plants are termed zoochorous; common examples include cocklebur, unicorn plant, and beggarticks, beggarticks (or Spanish needle). By developments of mutual evolution, the fleshy produce of fruits typically appeals to hungry animals, such that the seeds contained within are taken in, carried away, and later deposited (i.e., Defecation, defecated) at a distance from the parent plant. Likewise, the nutritious, oily kernels of Nut (fruit), nuts typically motivate birds and squirrels to hoarding, hoard them, burying them in soil to retrieve later during the winter of scarcity; thereby, uneaten seeds are sown effectively under natural conditions to Germination, germinate and grow a new plant some distance away from the parent. Other fruits have evolved flattened and elongated wings or helicopter-like blades, e.g.,
elm Elms are deciduous and semi-deciduous trees comprising the genus ''Ulmus'' in the family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical- montane regions of North America and Eurasia, ...
, maple, and tuliptree. This mechanism increases dispersal distance away from the parent via wind. Other wind-dispersed fruit have tiny "Pappus (flower structure), parachutes", e.g.,
dandelion ''Taraxacum'' () is a genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus has a near-cosmopolitan distribu ...
, Asclepias, milkweed, Tragopogon, salsify. Coconut fruits can float thousands of miles in the ocean, thereby spreading their seeds. Other fruits that can disperse via water are nipa palm and screw pine. Some fruits have evolved propulsive mechanisms that fling seeds substantial distances – perhaps up to in the case of the sandbox tree – via explosive dehiscence or other such mechanisms (see impatiens and squirting cucumber).


Food uses

A large variety of fruits – fleshy (simple) fruits from apples to berries to watermelon; dry (simple) fruits including beans and rice and coconuts; aggregate fruits including strawberries, raspberries, blackberries, pawpaw; and multiple fruits such as pineapple, fig, mulberries – are commercially valuable as human food. They are eaten both fresh and as jams, marmalade and other fruit preserves. They are used extensively in manufactured and processed foods (cakes, cookies, baked goods, flavorings, ice cream, yogurt, canned vegetables, frozen vegetables and meals) and beverages such as fruit juices and alcoholic beverages (brandy, fruit beer, wine). Spices like vanilla, black pepper, paprika, and
allspice Allspice, also known as Jamaica pepper, myrtle pepper, pimenta, or pimento, is the dried unripe berry of ''Pimenta dioica'', a midcanopy tree native to the Greater Antilles, southern Mexico, and Central America, now cultivated in many warm par ...
are derived from berries. Olive, Olive fruit is pressed for olive oil and similar processing is applied to other oil-bearing fruits and vegetables. Some fruits are available all year round, while others (such as blackberries and apricots in the UK) are subject to seasonal food, seasonal availability. Fruits are also used for socializing and gift-giving in the form of fruit baskets and fruit bouquets. Typically, many botanical fruits – "vegetables" in culinary parlance – (including tomato, green beans, leaf greens, bell pepper, cucumber, eggplant, okra, pumpkin, squash, zucchini) are bought and sold daily in fresh produce markets and greengroceries and carried back to kitchens, at home or restaurant, for preparation of meals.


Storage

All fruits benefit from proper post-harvest care, and in many fruits, the plant hormone Ethylene-ripened fruits, ethylene causes
ripening Ripening is a process in fruits that causes them to become more palatable. In general, fruit becomes sweeter, less green, and softer as it ripens. Even though the acidity of fruit increases as it ripens, the higher acidity level does not make t ...
. Therefore, maintaining most fruits in an efficient cold chain is optimal for post-harvest storage, with the aim of extending and ensuring shelf life.Why Cold Chain for Fruits:


Nutritional value

A meta-analysis of 83 studies showed fruit or vegetable consumption is associated with reduced markers of inflammation (reduced tumor necrosis factor and C-reactive protein) and enhanced immune cell profile (increased gamma delta T cells). Various culinary fruits provide significant amounts of Dietary fiber, fiber and water, and many are generally high in vitamin C. An overview of numerous studies showed that fruits (e.g., whole apples or whole oranges) are satisfying (filling) by simply eating and chewing them. The dietary fiber consumed in eating fruit promotes satiety, and may help to control body weight and aid reduction of blood cholesterol, a Risk factor (epidemiology), risk factor for cardiovascular diseases. Fruit consumption is under preliminary research for the potential to improve nutrition and affect chronic diseases. Regular consumption of fruit is generally associated with reduced risks of several diseases and functional declines associated with aging.


Food safety

For food safety, the Centers for Disease Control and Prevention, CDC recommends proper fruit handling and preparation to reduce the risk of food contamination and foodborne illness. Fresh fruits and vegetables should be carefully selected; at the store, they should not be damaged or bruised; and precut pieces should be refrigerated or surrounded by ice. All fruits and vegetables should be rinsed before eating. This recommendation also applies to produce with rinds or skins that are not eaten. It should be done just before preparing or eating to avoid premature spoilage. Fruits and vegetables should be kept separate from raw foods like meat, poultry, and seafood, as well as from utensils that have come in contact with raw foods. Fruits and vegetables that are not going to be cooked should be thrown away if they have touched raw meat, poultry, seafood, or eggs. All cut, peeled, or cooked fruits and vegetables should be refrigerated within two hours. After a certain time, harmful bacteria may grow on them and increase the risk of foodborne illness.


Allergies

Fruit allergies make up about 10 percent of all food-related allergies.


Nonfood uses

Because fruits have been such a major part of the human diet, various cultures have developed many different uses for fruits they do not depend on for food. For example: * Bayberry fruits provide a wax often used to make candles; * Many dry fruits are used as decorations or in dried flower arrangements (e.g., annual honesty, cotoneaster, Nelumbo, lotus,
milkweed ''Asclepias'' is a genus of herbaceous, perennial, flowering plants known as milkweeds, named for their latex, a milky substance containing cardiac glycosides termed cardenolides, exuded where cells are damaged. Most species are toxic to huma ...
, unicorn plant, and
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
). Ornamental trees and shrubs are often cultivated for their colorful fruits, including beautyberry, cotoneaster, holly, pyracantha, skimmia, and viburnum. * Fruits of opium poppy are the source of opium, which contains the drugs codeine and morphine, as well as the biologically inactive chemical theabaine from which the drug oxycodone is synthesized. * Osage orange fruits are used to repel cockroaches. * Many fruits provide natural dyes (e.g.,
cherry A cherry is the fruit of many plants of the genus ''Prunus'', and is a fleshy drupe (stone fruit). Commercial cherries are obtained from cultivars of several species, such as the sweet '' Prunus avium'' and the sour '' Prunus cerasus''. The na ...
, mulberry, sumac, and
walnut A walnut is the edible seed of any tree of the genus '' Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. They are accessory fruit because the outer covering of the fruit is technically an i ...
). * Dried gourds are used as bird houses, cups, decorations, dishes, musical instruments, and water jugs. * Pumpkins are carved into Jack-o'-lanterns for Halloween. * The fibrous core of the mature and dry Luffa fruit is used as a sponge. * The spiny fruit of burdock or cocklebur inspired the invention of Velcro. * Coir fiber from
coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family (biology), family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, ...
shells is used for brushes, doormats, floor tiles, insulation, mattresses, sacking, and as a growing medium for container plants. The shell of the coconut fruit is used to make bird houses, bowls, cups, musical instruments, and souvenir heads. * The hard and colorful grain fruits of Job's tears are used as decorative beads for jewelry, garments, and ritual objects. * Fruit is often a subject of still life paintings.


See also

* Fruit tree * Fruitarianism * List of countries by fruit production * List of culinary fruits * List of foods * List of fruit dishes


References


Further reading

* Gollner, Adam J. (2010). ''The Fruit Hunters: A Story of Nature, Adventure, Commerce, and Obsession''. Scribner. . * Watson, R. R., and Preedy, V.R. (2010, eds.). ''Bioactive Foods in Promoting Health: Fruits and Vegetables''. Academic Press. .


External links


Images of fruit development from flowers
at bioimages.Vanderbilt.edu (archived 18 February 2007)

at bioimages.Vanderbilt.edu (archived 25 April 2017)

( from California Rare Fruit Growers, Inc.)
Photo ID of Fruits
( by Capt. Pawanexh Kohli) * {{Authority control Fruit, Pollination Types of food